Difference between revisions of "Sapo National Park"

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[[West Africa]] > [[Liberia]] > [[Sapo National Park]]
 
[[West Africa]] > [[Liberia]] > [[Sapo National Park]]
  
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= Summary = <!-- An overview of the site, with one sentence for each section. May include a site map -->
+
__TOC__
<div style="float: right">
+
= Summary =
{{#display_map: height=200px | width=300px | scrollzoom=off | zoom=5 | layers= OpenStreetMap, OpenTopoMap
 
|5.378432, -8.496117~[[Sapo National Park]]~Western Chimpanzee
 
}}
 
</div>
 
  
 +
<div style="float: right">{{#display_map: height=190px | width=300px | scrollzoom=off | zoom=5 | layers= OpenStreetMap, OpenTopoMap|5.378432, -8.496117~[[Sapo National Park]]~'Pan troglodytes verus''}}</div>
 
* Western chimpanzees (''Pan troglodytes verus'') are present in Sapo National Park.
 
* Western chimpanzees (''Pan troglodytes verus'') are present in Sapo National Park.
 
* It has been estimated that 1,055 (CI: 595-1,870) individuals occur in the site.
 
* It has been estimated that 1,055 (CI: 595-1,870) individuals occur in the site.
Line 18: Line 12:
 
* The park has a total size of 1,804 km².
 
* The park has a total size of 1,804 km².
 
* Key threats to chimpanzees are poaching and illegal mining.
 
* Key threats to chimpanzees are poaching and illegal mining.
* Conservation activities have focused on long-term bio-monitoring and law enforcement.  
+
* Conservation activities have focused on long-term bio-monitoring and law enforcement.
 
* Sapo National Park is Liberia's first protected area, established in 1983.
 
* Sapo National Park is Liberia's first protected area, established in 1983.
  
= Site characteristics = <!-- A paragraph summary of physical and geographic aspects of the site, and a table of key information -->
 
  
Located in southeastern Liberia, Sapo National Park is Liberia's first protected area and represents one of the most intact forest ecosystems of the country (Tweh et al. 2018). The area of the park was extended from 1,304 km² to 1,804 km² in 2003 (Tweh et al. 2018). The park forms part of the Upper Guinean Forest ecosystem, and contains high levels of biodiversity (N'Goran et al. 2010). The park is a low elevation tropical humid rainforest. Elevation in the southeastern area is approximately 100 m with gently rolling hills while in the north, the elevation is approximately 400 m in the north with steep ridges (Peal & Kranz 1990). In addition to the western chimpanzee, other endangered and vulnerable species inhabit the site, including forest elephants (''Loxodonta africana''), pygmy hippopotamus (''Hexaprotodon liberiensis''), Jentink’s duiker (''Cephalophus jentinki''), red colobus (''Piliocolobus badius''), and Diana monkeys (''Cercopithecus diana diana'', N'Goran 2010).  
+
 
 +
= Site characteristics =
 +
 
 +
Located in southeastern Liberia, Sapo National Park is Liberia's first protected area and represents one of the most intact forest ecosystems of the country (Tweh et al. 2018). The area of the park was extended from 1,304 km² to 1,804 km² in 2003 (Tweh et al. 2018). The park forms part of the Upper Guinean Forest ecosystem, and contains high levels of biodiversity (N'Goran et al. 2010). The park is a low elevation tropical humid rainforest. Elevation in the southeastern area is approximately 100m with gently rolling hills while in the north, the elevation is approximately 400m in the north with steep ridges (Peal & Kranz 1990). In addition to the western chimpanzee, other endangered and vulnerable species inhabit the site, including forest elephants (''Loxodonta africana''), pygmy hippopotamus (''Hexaprotodon liberiensis''), Jentink’s duiker (''Cephalophus jentinki''), red colobus (''Piliocolobus badius''), and Diana monkeys (''Cercopithecus diana diana'', N'Goran 2010).
  
 
'''Table 1. Basic site information for Sapo National Park'''
 
'''Table 1. Basic site information for Sapo National Park'''
{| border="1" cellpadding="5" cellspacing="0" class="basic-information"
+
{| border="1" cellpadding="5" cellspacing="0" class="Site_characteristics-table"
| Species
+
|Species
|''Pan troglodytes verus''
+
|'Pan troglodytes verus''
 
|-
 
|-
| Area
+
|Area
|1804 km²
+
|1,804 km²
 
|-
 
|-
 
|Coordinates
 
|Coordinates
|5.378432, -8.496117
+
|Lat: 5.378432 , Lon:  -8.496117
 
|-
 
|-
 
|Type of site
 
|Type of site
 
|Protected area (National Park)
 
|Protected area (National Park)
 
|-
 
|-
|Governance type
+
|Habitat types
|Governance by government
+
|Subtropical/tropical moist lowland
 
|-
 
|-
|Habitat type
+
|Type of governance
|Subtropical/tropical moist lowland
+
|
 
|}
 
|}
[[Types of sites]] ⋅ [[Governance types]] ⋅ [[Habitat types]]
 
  
= Ape status = <!--An overview of ape population status (population sizes, trends, etc.), followed by a table of specific surveys and results -->
+
[https://www.iucnredlist.org/resources/habitat-classification-scheme IUCN habitat categories] [[Site designations]]
  
A survey in 1982 (one year before the establishment of the park), confirmed the presence of chimpanzees in the Sapo forest (Anderson et al. 1983). Based on the estimates from two surveys, one in 2009 (N'Goran et al. 2010) and a second one in 2017 (Tweh et al. 2018), the chimpanzee population in the park has remained relatively stable, with an estimated abundance of approximately 1,055 individuals.
+
= Ape status =
  
'''Table 2. Ape population estimates in Sapo National Park'''
+
A survey in 1982 (one year before the establishment of the park), confirmed the presence of chimpanzees in the Sapo forest (Anderson et al. 1983). Based on the estimates from two surveys, one in 2009 (N'Goran et al. 2010) and a second one in 2017 (Tweh et al. 2018), the chimpanzee population in the park has remained relatively stable, with an estimated abundance of approximately 1,055 individuals.
{| border="1" cellpadding="5" cellspacing="0" class="population-estimate-table"
+
 
! Species
+
'''Table 2. Ape population estimates reported for Sapo National Park'''
! Year
+
{| border="1" cellpadding="5" cellspacing="0" class="Ape_status-table"
! Occurrence
+
!Species
! Encounter or visitation rate (nests/km; ind/day)
+
!Year
! Density estimate [ind/ km²] (95% CI)
+
!Occurrence
! Abundance estimate (95% CI)
+
!Encounter or vistation rate (nests/km; ind/day)
! Survey area
+
!Density estimate [ind./ km²] (95% CI)
! Sampling method
+
!Abundance estimate (95% CI)
! Analytical framework
+
!Survey area
! Source
+
!Sampling method
! Comments
+
!Analytical framework
! A.P.E.S. database ID
+
!Source
 +
!Comments
 +
!A.P.E.S. database ID
 
|-
 
|-
 
|''Pan troglodytes verus''
 
|''Pan troglodytes verus''
Line 72: Line 69:
 
|0.24
 
|0.24
 
|
 
|
|Southeastern sector (50 km²)
+
|Southeastern sector of the park (50 km²)
 
|Line transects
 
|Line transects
 
|
 
|
Line 94: Line 91:
 
|''Pan troglodytes verus''
 
|''Pan troglodytes verus''
 
|2007-2009
 
|2007-2009
|Present
+
|
 
|0.27
 
|0.27
 
|
 
|
 
|
 
|
|Sapo National Park, excluding southeast area
+
|Sapo National Park, excluding south east area
 
|Line transects
 
|Line transects
 
|
 
|
Line 107: Line 104:
 
|''Pan troglodytes verus''
 
|''Pan troglodytes verus''
 
|2009
 
|2009
|Present
+
|
 
|4.05
 
|4.05
 
|0.86
 
|0.86
|1079 (713-1633)
+
|1079 (CI: 713-1633)
 
|Sapo National Park, excluding mining areas
 
|Sapo National Park, excluding mining areas
 
|Line transects
 
|Line transects
Line 120: Line 117:
 
|''Pan troglodytes verus''
 
|''Pan troglodytes verus''
 
|2016-2017
 
|2016-2017
|Present
 
 
|
 
|
|0.83  
+
|
|1055 (595-1870)
+
|0.83
|Sapo National Park, excluding southeast area
+
|1055 (CI: 595-1870)
 +
|Sapo National Park, excluding south east area
 
|Line transects
 
|Line transects
 
|
 
|
Line 131: Line 128:
 
|
 
|
 
|}
 
|}
[[Sampling methods]] ⋅ [[Analytical frameworks]]
 
  
= Threats =    <!-- a text overview of threats, followed by a table of key threats -->
 
  
Sapo National Park has been primarily threatened by illegal hunting and mining (Tweh et al. 2018; Greengrass 2015; N'Goran et al. 2010). An estimated 18,000 illegal miners were inhabiting the park in 2010, the majority of which was evicted by the government in the same year (Vogt 2011). A survey of two commercial hunting camps bordering the park revealed high hunting pressure in the area, and the majority of bushmeat harvested was destined to urban areas (Greengrass 2015). The carcasses documented during this survey included chimpanzees as well as other endangered and vulnerable species, such as the red colobus monkey, Diana monkey, and pygmy hippopotamus. Furthermore, the development of the road network around the park is expected to increase hunting pressure and facilitate the bushmeat trade (Greengrass 2015), as well as other illegal activities in the park.
+
= Threats =
  
<div><ul>
+
Sapo National Park has been primarily threatened by illegal hunting and mining (Tweh et al. 2018; Greengrass 2015; N'Goran et al. 2010). An estimated 18,000 illegal miners were inhabiting the park in 2010, the majority of which was evicted by the government on the same year (Vogt 2011). A survey of two commercial hunting camps bordering the park revealed high hunting pressure in the area, and the majority of bushmeat harvested was destined to urban areas (Greengrass 2015). The carcasses documented during this survey included chimpanzees as well as other endangered and vulnerable species, such as the red colobus monkey, Diana monkey, and pygmy hippopotamus. Furthermore, the development of the road network around the park is expected to increase hunting pressure and facilitate the bushmeat trade (Greengrass 2015), as well as other illegal activities in the park.
<li style="display: inline-block; vertical-align: top;"> [[File: IllegalMining_Sapo_BTally.jpg | 200px | thumb| right | Illegal mining activity in Sapo © Ben Tally]] </li>
 
<li style="display: inline-block; vertical-align: top;"> [[File: MiningCamp_Sapo_BTally.jpg | 200px | thumb| right | Mining camp © Ben Tally]] </li>
 
<li style="display: inline-block; vertical-align: top;"> [[File: IllegalMining_BTally.jpg | 200px | thumb| right |Illegal mining © Ben Tally]] </li>
 
<li style="display: inline-block; vertical-align: top;"> [[File: Patrol_SMART.jpg | 200px | thumb| right |Illegal mining © Ben Tally]] </li>
 
</ul></div>
 
  
'''Table 3. Threats to apes in Sapo National Park'''
+
'''Table 3. Threats to apes reported for Sapo National Park'''
{| border="1" cellpadding="5" cellspacing="0" class="threats-table"
+
{| border="1" cellpadding="5" cellspacing="0" class="Threats-table"
!align="left"|Category <!-- PLEASE DO NOT CHANGE THE THREAT CATEGORIES -->
+
!Category
 
!Specific threats
 
!Specific threats
 
!Threat level
 
!Threat level
Line 152: Line 142:
 
!Year of threat
 
!Year of threat
 
|-
 
|-
|1. Residential & commercial development
+
|10 Geological events
|1.1 Housing & urban areas
+
|
|High  
+
|Absent
|Illegal settlement of miners; 18,000 settlers in 2010 (Vogt 2011).
+
|
 +
|
 +
|-
 +
|12 Other threat
 +
|
 +
|Absent
 +
|
 +
|
 +
|-
 +
|1 Residential & commercial development
 +
|1.1 Residential areas
 +
|High (more than 70% of population affected)
 +
|Illegal settlement of miners (Vogt 2011).
 
|2010
 
|2010
 
|-
 
|-
|2. Agriculture & aquaculture
+
|3 Energy production & mining
|2.1 Annual & perennial non-timber crops
+
|3.2.3 Artisanal mining
|Present (severity unknown)
+
|High (more than 70% of population affected)
|Present as a result of illegal settlements; scale unknown (N’Goran et al. 2010)
+
|Illicit gold mining which has decreased since 2010 (Tweh et al. 2018); artisanal mining is still present (Junket per. comm. 2019).
|2010-Ongoing (2023)
+
|Ongoing ( 2019)
 
|-
 
|-
|3. Energy production & mining
+
|5 Biological resource use
|3.2.3 Artisanal mining
+
|5.1 Hunting & collecting terrestrial animals
|High  
+
|High (more than 70% of population affected)
|Illicit gold mining which has decreased since 2010 (Tweh et al. 2018); artisanal mining is still present (Junker personal. comm. 2019).
+
|Poaching represents a major threat to chimpanzees and other species in the park (Tweh et al. 2018, N’Goran et al. 2010, Greengrass 2015), and most of the bushmeat is destined to urban areas (Greengrass 2015).
|2010-Ongoing (2019)
+
|Ongoing (2010, 2015, 2018)
 
|-
 
|-
|4. Transportation & service corridors
+
|6 Human intrusions & disturbance
|4.1 Roads & railroads
+
|6.2 War, civil unrest & military exercises
|Present (severity unknown)
+
|High (more than 70% of population affected)
|Development of the road network around the park facilitates illegal human activities in the park (Greengrass 2015).
+
|Two civil wars since the establishment of the park disrupted conservation activities, and led to illegal occupation of the park, as well as poaching and extraction of natural resources (Greengrass 2015, Collen et al. 2011).
|2015-Ongoing (2023)
+
|1989-1996, 1999-2003
 
|-
 
|-
|5. Biological resource use
+
|2 Agriculture & aquaculture
|5.1.3 Commercial bushmeat trade
+
|2.1 Annual & perennial non-timber crops
|High
+
|Present (unknown severity)
|Poaching represents a major threat to chimpanzees and other species in the park (Tweh et al. 2018, N’Goran et al. 2010, Greengrass 2015), and most of the bushmeat is destined to urban areas (Greengrass 2015). Hunting sign encounter rate: 1.7/km (Tweh et al. 2018).
+
|Present as a result of illegal settlements; scale unknown (N’Goran et al. 2010).
|2010-Ongoing (2023)
+
|Ongoing (2010)
 
|-
 
|-
|6. Human intrusion & disturbance
+
|4 Transportation & service corridors
|6.2 War, civil unrest & military exercises
+
|4.1 Roads & railroads
|High
+
|Present (unknown severity)
|Two civil wars since the establishment of the park disrupted conservation activities, and led to illegal occupation of the park, as well as poaching and extraction of natural resources (Greengrass 2015, Collen et al. 2011)
+
|Development of the road network around the park facilitates illegal human activities in the park (Greengrass 2015).
|1989-1996, 1999-2003
+
|Ongoing (2015)
 
|-
 
|-
|7. Natural system modifications
+
|7 Natural system modifications
 
|
 
|
 
|Unknown
 
|Unknown
Line 194: Line 196:
 
|
 
|
 
|-
 
|-
|8. Invasive & other problematic species, genes, diseases
+
|8 Invasive & other problematic species, genes & diseases
 
|
 
|
 
|Unknown
 
|Unknown
Line 200: Line 202:
 
|
 
|
 
|-
 
|-
|9. Pollution
+
|9 Pollution
 
|
 
|
 
|Unknown
 
|Unknown
Line 206: Line 208:
 
|
 
|
 
|-
 
|-
|10. Geological Events
+
|11 Climate change & severe weather
|
 
|Absent
 
|
 
|
 
|-
 
| 11. Climate change & severe weather
 
 
|
 
|
 
|Unknown
 
|Unknown
 
|
 
|
 
|
 
|
|-
 
|12. Other options
 
|
 
|Absent
 
|
 
|
 
|-
 
 
|}
 
|}
[[Threats]]
 
  
= Conservation activities =       <!-- A summary of the conservation activities, followed by a table of key activities -->
+
[https://www.iucnredlist.org/resources/threat-classification-scheme IUCN Threats list]
 +
<div><ul><li style="display: inline-block; vertical-align: top;"> [[File: IllegalMining_Sapo_BTally.jpg | 200px | thumb| right | Illegal mining activity in Sapo © Ben Tally]] </li><li style="display: inline-block; vertical-align: top;"> [[File: MiningCamp_Sapo_BTally.jpg | 200px | thumb| right | Mining camp © Ben Tally]] </li><li style="display: inline-block; vertical-align: top;"> [[File: IllegalMining_BTally.jpg | 200px | thumb| right |Illegal mining © Ben Tally]] </li><li style="display: inline-block; vertical-align: top;"> [[File: Patrol_SMART.jpg | 200px | thumb| right |Illegal mining © Ben Tally]] </li></ul></div>
 +
= Conservation activities =
  
 
The Forestry Development Authority of Liberia is responsible for the sustainable management of the forest sector and the protection of all natural resources. It runs the Sapo National Park in collaboration with Fauna & Flora International (FFI) and Wild Chimpanzee Foundation (WCF).  Main activities at the national park level include anti-poaching, conservation education awareness, and bio-monitoring & scientific research. As of 2019, the main activities in the research area have been camera trappings (2019,2020, 2021) by FFI and FDA in the entire national park. eDNA (focusing on Pygmy hippopotamus) was conducted in 2022. The national park forms part of the Tai-Grebo-Sapo Forest Complex, which is a conservation priority in West Africa. Conservation efforts in Sapo National Park have mainly focused on law enforcement, conservation awareness, and bio-monitoring. The WCF has supported Community Watch Teams (CWT), which comprise members from surrounding communities, and regularly patrol and support FDA rangers (WCF 2019). CWTs have played an important role in the eviction of illegal miners from the national park (WCF 2019). In 2012, Fauna & Flora International established a long-term bio-monitoring program to follow the population trends for chimpanzees, pygmy hippopotamuses, elephants, as well as duikers, birds, reptiles, and amphibians (Tweh et al. 2018). Together with Liberia's Forestry Development Authority, permanent transects are surveyed twice a year as part of this long-term bio-monitoring program (Tweh et al. 2018).
 
The Forestry Development Authority of Liberia is responsible for the sustainable management of the forest sector and the protection of all natural resources. It runs the Sapo National Park in collaboration with Fauna & Flora International (FFI) and Wild Chimpanzee Foundation (WCF).  Main activities at the national park level include anti-poaching, conservation education awareness, and bio-monitoring & scientific research. As of 2019, the main activities in the research area have been camera trappings (2019,2020, 2021) by FFI and FDA in the entire national park. eDNA (focusing on Pygmy hippopotamus) was conducted in 2022. The national park forms part of the Tai-Grebo-Sapo Forest Complex, which is a conservation priority in West Africa. Conservation efforts in Sapo National Park have mainly focused on law enforcement, conservation awareness, and bio-monitoring. The WCF has supported Community Watch Teams (CWT), which comprise members from surrounding communities, and regularly patrol and support FDA rangers (WCF 2019). CWTs have played an important role in the eviction of illegal miners from the national park (WCF 2019). In 2012, Fauna & Flora International established a long-term bio-monitoring program to follow the population trends for chimpanzees, pygmy hippopotamuses, elephants, as well as duikers, birds, reptiles, and amphibians (Tweh et al. 2018). Together with Liberia's Forestry Development Authority, permanent transects are surveyed twice a year as part of this long-term bio-monitoring program (Tweh et al. 2018).
  
'''Table 4. Conservation activities in Sapo National Park'''
+
'''Table 4. Conservation activities reported for Sapo National Park'''
{| border="1" cellpadding="5" cellspacing="0" class="conservation-actions-table"
+
{| border="1" cellpadding="5" cellspacing="0" class="Conservation_activities-table"
!align="left"|Category <!-- PLEASE DO NOT CHANGE THE CATEGORIES -->
+
!Category
 
!Specific activity
 
!Specific activity
!Description  
+
!Description
!Implementing organization
+
!Implementing organization(s)
 
!Year of activity
 
!Year of activity
 
|-
 
|-
|1. Development impact mitigation
+
|2 Counter-wildlife crime
|Not reported
+
|2.3 Conduct regular anti-poaching patrols
|
+
|Community Watch Teams supported by the WCF regularly patrol the site (WCF 2019)
|
 
 
|
 
|
 +
|Ongoing (2019)
 
|-
 
|-
|2. Counter-wildlife crime
+
|2 Counter-wildlife crime
|2.3 Conduct regular anti-poaching patrols
 
|Community Watch Teams supported by the WCF regularly patrol the site (WCF 2019).
 
|Wild Chimpanzee Foundation
 
|2019-Ongoing (2023)
 
|-
 
|2. Counter-wildlife crime
 
 
|2.8 Provide training to anti-poaching ranger patrols
 
|2.8 Provide training to anti-poaching ranger patrols
|Members of the Community Watch Teams are trained in the use of equipment (GPS, compass, camera) and patrolling (WCF 2019).
+
|Members of the Community Watch Teams are trained in the use of equipment (GPS, compass, camera) and patrolling (WCF 2019)
|Wild Chimpanzee Foundation
+
|
|2019-Ongoing (2023)
+
|Ongoing (2019)
 
|-
 
|-
|2. Counter-wildlife crime
+
|2 Counter-wildlife crime
 
|2.11 Implement monitoring surveillance strategies (e.g., SMART) or use monitoring data to improve effectiveness of patrols
 
|2.11 Implement monitoring surveillance strategies (e.g., SMART) or use monitoring data to improve effectiveness of patrols
 
|Long-term bio-monitoring of chimpanzee population (Tweh et al. 2018)
 
|Long-term bio-monitoring of chimpanzee population (Tweh et al. 2018)
 
|
 
|
|2019-Ongoing (2023)
+
|Ongoing (2019)
 
|-
 
|-
|3. Species health
+
|4 Education & awareness
|Not reported
 
|
 
|
 
|
 
|-
 
|4. Education & awareness
 
 
|4.2 Involve local community in ape research and conservation management
 
|4.2 Involve local community in ape research and conservation management
|As part of a long-term bio-monitoring program, staff of Liberia’s Forestry Development Authority and members of the local community are involved in the surveys (Tweh et al. 2018).
+
|As part of a long-term bio-monitoring program, staff of Liberia’s Forestry Development Authority and members of the local community are involved in the surveys (Tweh et al. 2018)
|
 
|2018-Ongoing (2023)
 
|-
 
|5. Protection & restoration
 
|5.2 Legally protect ape habitat
 
|The area is designated as a National Park.
 
 
|
 
|
|1983-Ongoing (2023)
+
|Ongoing (2018)
 
|-
 
|-
|5. Protection & restoration
+
|5 Protection & restoration
 
|5.9 Resettle illegal human communities (i.e., in a protected area) to another location
 
|5.9 Resettle illegal human communities (i.e., in a protected area) to another location
|Eviction of 18,000 illegal settlers in the park by the Liberia's government (Vogt 2011).
+
|Eviction of 18,000 illegal settlers in the park by the Liberia's government (Vogt 2011)
 
|
 
|
 
|2010
 
|2010
 
|-
 
|-
|6. Species management
+
|5 Protection & restoration
|Not reported
+
|5.2 Legally protect ape habitat
 +
|The area is designated as National Park (N’Goran et al. 2010)
 
|
 
|
|
+
|Ongoing (2010)
|
 
|-
 
|7. Economic & other incentives
 
|Not reported
 
|
 
|
 
|
 
|-
 
|8. Permanent presence
 
|Not reported
 
|
 
|
 
|
 
|-
 
 
|}
 
|}
[[Conservation activities]]
 
  
= Conservation implementation challenges and enablers = <!-- Overview of challenges in ape conservation -->
+
[[Conservation activities list (Junker et al. 2017)]]
 +
 
 +
= Challenges =
  
 
The influx of local community dwellers inside the national park is hampering the SNP management to implement the protection of chimpanzees. Low manpower for conducting anti-poaching patrols in and outside the park has put chimpanzees under serious threat.
 
The influx of local community dwellers inside the national park is hampering the SNP management to implement the protection of chimpanzees. Low manpower for conducting anti-poaching patrols in and outside the park has put chimpanzees under serious threat.
Very high illiteracy among the rangers thus making it difficult to collect data, process court procedures, and conduct conservation education awareness to local community people.
 
  
 
'''Table 5. Challenges reported for Sapo National Park'''
 
'''Table 5. Challenges reported for Sapo National Park'''
{| border="1" cellpadding="5" cellspacing="0" class="challenges-table"
+
{| border="1" cellpadding="5" cellspacing="0" class="Challenges-table"
!align="left"|Category <!-- PLEASE DO NOT CHANGE THE CATEGORIES -->
+
!Challenges
!Challenge
+
!Specific challenges
 
!Source
 
!Source
 
!Year(s)
 
!Year(s)
 
|-
 
|-
|1. Site management
+
|4 Institutional support
|1.4 Conflict on land tenure
+
|4.1 Lack of law enforcement
|Tally, B. pers. comm. 2022
+
|Greengrass 2015, N'Goran et al. 2010
|Unknown
+
|
 
|-
 
|-
|2. Resources & capacity
+
|2 Resources and capacity
 
|2.5 Lack of equipment/transportation
 
|2.5 Lack of equipment/transportation
|Waitkuwait 2003
+
|Tweh et al. 2018
|Unknown
 
|-
 
|2. Resources & capacity
 
|2.2 Lack of staff
 
|Tally, B. pers. comm. 2022
 
|Unknown
 
|-
 
|3. Engaged community
 
|Not reported
 
|
 
 
|
 
|
 
|-
 
|-
|4. Institutional support
+
|1 Site management
|4.1 Lack of law enforcement
+
|1.2 Need for improved coordination
|Greengrass 2015, N'Goran et al. 2010
+
|Waitkuwait 2003
|Unknown
 
|-
 
|5. Ecological context
 
|Not reported
 
|
 
 
|
 
|
 
|-
 
|-
|6. Safety & stability
+
|1 Site management
|Not reported
+
|1.4 Conflict on land tenure
 +
|Tally, B. pers. comm. 2022
 
|
 
|
|
 
|-
 
 
|}
 
|}
[[Challenges]]
+
 
 +
 
 +
= Enablers =
 +
 
 +
 
  
 
'''Table 6. Enablers reported for Sapo National Park'''
 
'''Table 6. Enablers reported for Sapo National Park'''
{| border="1" cellpadding="5" cellspacing="0" class="enablers-table"
+
{| border="1" cellpadding="5" cellspacing="0" class="enabler-table"
!align="left"|Category <!-- PLEASE DO NOT CHANGE THE CATEGORIES -->
+
!Enablers
!Enabler
+
!Specific enablers
 
!Source
 
!Source
 
!Year(s)
 
!Year(s)
 
|-
 
|-
|1. Site management
+
|1 Site management
|Not reported
+
|
 
|
 
|
 
|
 
|
 
|-
 
|-
|2. Resources & capacity
+
|2 Resources and capacity
|Not reported
+
|
 
|
 
|
 
|
 
|
 
|-
 
|-
|3. Engaged community
+
|3 Engaged community
|Not reported
+
|
 
|
 
|
 
|
 
|
 
|-
 
|-
|4. Institutional support
+
|4 Institutional support
|Not reported
+
|
 
|
 
|
 
|
 
|
 
|-
 
|-
|5. Ecological context
+
|5 Ecological context
|Not reported
+
|
 
|
 
|
 
|
 
|
 
|-
 
|-
|6. Safety & stability
+
|6 Safety and stability
|Not reported
+
|
 
|
 
|
 
|
 
|
|-
 
 
|}
 
|}
[[Enablers]]
 
  
= Research activities = <!-- Overview of research activities -->
+
 
 +
= Research activities =
  
 
Several surveys have been done in the park to monitor the chimpanzee population (e.g., N'Goran et al. 2010, Tweh et al. 2018), assess the impacts of conservation interventions (Tweh et al. 2018), investigate the behavior and ecology of chimpanzees in the park (Anderson et al. 1983), and investigate the impact of hunting pressure in the area (Greengrass 2015).
 
Several surveys have been done in the park to monitor the chimpanzee population (e.g., N'Goran et al. 2010, Tweh et al. 2018), assess the impacts of conservation interventions (Tweh et al. 2018), investigate the behavior and ecology of chimpanzees in the park (Anderson et al. 1983), and investigate the impact of hunting pressure in the area (Greengrass 2015).
  
===Documented behaviours===  <!-- List of any behaviours observed at the site, including citations -->
 
  
'''Table 7. Ape behaviors reported for Sapo National Park'''
+
= Documented behaviours =
{| border="1" cellpadding="5" cellspacing="0" class="behaviors-table"
+
 
!align="left"|Behavior
+
 
 +
 
 +
'''Table 7. Behaviours documented for Sapo National Park'''
 +
{| border="1" cellpadding="5" cellspacing="0" class="behaviours-table"
 +
!Behavior
 
!Source
 
!Source
 
|-
 
|-
 
|Nut cracking
 
|Nut cracking
 
|Anderson et al. 1983
 
|Anderson et al. 1983
|-
 
 
|}
 
|}
 +
  
 
= Exposure to climate change impacts =
 
= Exposure to climate change impacts =
  
As part of a study on the exposure of African great ape sites to climate change impacts, Kiribou et al. (2024) extracted climate data and data on projected extreme climate impact events for the site. Climatological characteristics were derived from observation-based climate data provided by the Inter-Sectoral Impact Model Intercomparison Project ([ISIMIP www.isimip.org]). Parameters were calculated as the average across each 30-year period.
 
For future projections, two Representative Concentration Pathways (RCP) were used. RCP 2.6 is a scenario with strong mitigation measures in which global temperatures would likely rise below 2°C. RCP 6.0 is a scenario with medium emissions in which global temperatures would likely rise up to 3°C by 2100. For the number of days with heavy precipitation events, the 98th percentile of all precipitation days (>1mm/d) was calculated for the 1979-2013 reference period as a threshold for a heavy precipitation event. Then, for each year, the number of days above that threshold was derived. The figures on temperature and precipitation anomaly show the deviation from the mean temperature and mean precipitation for the 1979-2013 reference period. The estimated exposure to future extreme climate impact events (crop failure, drought, river flood, wildfire, tropical cyclone, and heatwave) is based on a published dataset by Lange et al. 2020 derived from ISIMIP2b data. The same global climate models and RCPs as described above were used. Within each 30-year period, the number of years with an extreme event and the average proportion of the site affected were calculated (Kiribou et al. 2024).
 
  
'''Table 8. Estimated past and projected climatological characteristics in Sapo National Park'''
 
{| border='1' cellpadding='5' cellspacing='0' class='EE-table'
 
|
 
|'''1981-2010'''
 
|'''2021-2050, RCP 2.6'''
 
|'''2021-2050, RCP 6.0'''
 
|'''2071-2099, RCP 2.6'''
 
|'''2071-2099, RCP 6.0'''
 
|-
 
|Mean temperature [°C]
 
|25.9
 
|26.9
 
|26.8
 
|26.9
 
|28
 
|-
 
|Annual precipitation [mm]
 
|2893
 
|2882
 
|3001
 
|2921
 
|2958
 
|-
 
|Max no. consecutive dry days (per year)
 
|17.5
 
|21
 
|16.9
 
|20.4
 
|19.1
 
|-
 
|No. days with heavy precipitation (per year)
 
|5.9
 
|12.5
 
|14.3
 
|14.1
 
|15.6
 
|-
 
|}
 
  
  
'''Table 9. Projected exposure of apes to extreme climate impact events in Sapo National Park'''
+
 
{| border='1' cellpadding='5' cellspacing='0' class='EE-table'
+
= External links =
|
+
 
|'''No. of years with event (2021-2050, RCP 2.6)'''
+
 
|'''% of site exposed (2021-2050, RCP 2.6)'''
+
 
|'''No. of years with event (2021-2050, RCP 6.0)'''
+
= Relevant datasets =
|'''% of site exposed (2021-2050, RCP 6.0)'''
 
|'''No. of years with event (2070-2099, RCP 2.6)'''
 
|'''% of site exposed (2070-2099, RCP 2.6)'''
 
|'''No. of years with event (2070-2099, RCP 6.0)'''
 
|'''% of site exposed (2070-2099, RCP 6.0)'''
 
|-
 
|Crop failure
 
|2
 
|0.04
 
|1
 
|0.04
 
|1
 
|0.04
 
|6
 
|0.04
 
|-
 
|Drought
 
|0
 
|0
 
|0
 
|0
 
|0
 
|0
 
|0
 
|0
 
|-
 
|Heatwave
 
|13
 
|100
 
|10.5
 
|100
 
|17.5
 
|100
 
|15
 
|100
 
|-
 
|River flood
 
|0
 
|0
 
|1
 
|4.2
 
|0.5
 
|0.92
 
|0.5
 
|0.08
 
|-
 
|Tropical cyclone
 
|0
 
|0
 
|1
 
|1.36
 
|0
 
|0
 
|0.5
 
|1
 
|-
 
|Wildfire
 
|30
 
|0.44
 
|30
 
|0.38
 
|29
 
|0.46
 
|29
 
|0.41
 
|-
 
|}
 
  
  
<div><ul>
 
<li style="display: inline-block; vertical-align: top;"> [[File: PrecipAnomaly Sapo NP.png | 450px | thumb| right | Precipitation anomaly in Sapo National Park]] </li>
 
<li style="display: inline-block; vertical-align: top;"> [[File: TempAnomaly Sapo NP.png | 450px | thumb| right | Temperature anomaly in Sapo National Park]] </li>
 
</ul></div>
 
  
 
= References =
 
= References =
Kiribou, R., Tehoda, P., Chukwu, O., Bempah, G., Kühl, H. S., Ferreira, J., ... & Heinicke, S. (2024). Exposure of African ape sites to climate change impacts. PLOS Climate, 3(2), e0000345.<br>
+
 
Lange, S., Volkholz, J., Geiger, T., Zhao, F., Vega, I., Veldkamp, T., ... & Frieler, K. (2020). Projecting exposure to extreme climate impact events across six event categories and three spatial scales. Earth's Future, 8(12), e2020EF001616.<br>
+
Tweh, C., Kouakou, C.Y., Chira, R., Freeman, B., Githaiga, J.M., Kerwillain, S., Molokwu-Odozi, M., Varney M. and Junker, J. 2018. Nest counts reveal a stable chimpanzee population in Sapo National Park, Liberia. Primate Conservation 2018 (32): 12 pp.
Tweh, C., Kouakou, C.Y., Chira, R., Freeman, B., Githaiga, J.M., Kerwillain, S., Molokwu-Odozi, M., Varney M. and Junker, J. 2018. Nest counts reveal a stable chimpanzee population in Sapo National Park, Liberia. Primate Conservation 2018 (32): 12 pp. <br>
+
 
N’Goran, K. P., Kouakou, C.Y. and Herbinger I. 2010. Report on the Population Survey and Monitoring of Chimpanzee in Sapo National Park, Liberia (June–December 2009). Report. Wild Chimpanzee Foundation, Abidjan, Côted’Ivoire. <br>
+
N’Goran, K. P., Kouakou, C.Y. and Herbinger I. 2010. Report on the Population Survey and Monitoring of Chimpanzee in Sapo National Park, Liberia (June–December 2009). Report. Wild Chimpanzee Foundation, Abidjan, Côted’Ivoire.
Anderson, R., Williamson, E.A., and Carter, J. 1983. Chimpanzees of Sapo Forest, Liberia: density, nests, tools and meat-eating. PRIMAaXS, 24(4): 594-601. <br>
+
 
Waitkuwait, W.E. 2003. Report on the First Year of Operation of a Community-based Bio-monitoring Programme in and around Sapo National Park, Sinoe County, Liberia. Report. Fauna and Flora International. <br>
+
Anderson, R., Williamson, E.A., and Carter, J. 1983. Chimpanzees of Sapo Forest, Liberia: density, nests, tools and meat-eating. PRIMAaXS, 24(4): 594-601.
Vogt, M. 2011. Results of Sapo National Park Bio-Monitoring Programme 2007-2009. Report. Fauna & Flora International, Monrovia, Liberia. <br>
+
 
Greengrass, E. 2015. Commercial hunting to supply urban markets threatens mammalian biodiversity in Sapo National Park. Oryx 50(3), 397–404. <br>
+
Waitkuwait, W.E. 2003. Report on the First Year of Operation of a Community-based Bio-monitoring Programme in and around Sapo National Park, Sinoe County, Liberia. Report. Fauna and Flora International.
Collen, B., Howard, B., Konie, J., Daniel, O., and Rist, J. 2011. Field surveys for the endangered pygmy hippopotamus Choerpsis liberiensis in Sapo National Park, Liberia. Oryx, 45(1), 35–37. <br>
+
 
Wild Chimpanzee Foundation. 2019. Activities of the Wild Chimpanzee Foundation for improved conservation of chimpanzees and their habitat in West Africa. Annual Report 2018.  
+
Vogt, M. 2011. Results of Sapo National Park Bio-Monitoring Programme 2007-2009. Report. Fauna & Flora International, Monrovia, Liberia.
Peal, A. L., & Kranz, K. R. (1990). Antelopes: GLobal Survey and Regional Action Plans, Part 3. West and  
+
 
 +
Greengrass, E. 2015. Commercial hunting to supply urban markets threatens mammalian biodiversity in Sapo National Park. Oryx 50(3), 397–404.
 +
 
 +
Collen, B., Howard, B., Konie, J., Daniel, O., and Rist, J. 2011. Field surveys for the endangered pygmy hippopotamus Choerpsis liberiensis in Sapo National Park, Liberia. Oryx, 45(1), 35–37.
 +
 
 +
Wild Chimpanzee Foundation. 2019. Activities of the Wild Chimpanzee Foundation for improved conservation of chimpanzees and their habitat in West Africa. Annual Report 2018.
 +
 
 +
Peal, A. L., & Kranz, K. R. (1990). Antelopes: GLobal Survey and Regional Action Plans, Part 3. West and
 +
 
 
Central Africa. Gland, Switzerland: World Conservation Union.
 
Central Africa. Gland, Switzerland: World Conservation Union.
  
<br>
+
 
'''Page completed by:''' Ben Tally & A.P.E.S. Wiki Team '''Date:''' 23/01/23  <!-- If you don't want to add your name, you can add "A.P.E.S. Wiki team" -->
+
 
 +
'''Page created by: '''Ben Tally & A.P.E.S. Wiki Team''' Date:''' NA

Latest revision as of 09:57, 18 March 2025

West Africa > Liberia > Sapo National Park

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Summary

Leaflet | Map data: © OpenStreetMap, SRTM | Map style: © OpenTopoMap (CC-BY-SA), © OpenStreetMap
  • Western chimpanzees (Pan troglodytes verus) are present in Sapo National Park.
  • It has been estimated that 1,055 (CI: 595-1,870) individuals occur in the site.
  • The chimpanzee population trend is stable.
  • The park has a total size of 1,804 km².
  • Key threats to chimpanzees are poaching and illegal mining.
  • Conservation activities have focused on long-term bio-monitoring and law enforcement.
  • Sapo National Park is Liberia's first protected area, established in 1983.


Site characteristics

Located in southeastern Liberia, Sapo National Park is Liberia's first protected area and represents one of the most intact forest ecosystems of the country (Tweh et al. 2018). The area of the park was extended from 1,304 km² to 1,804 km² in 2003 (Tweh et al. 2018). The park forms part of the Upper Guinean Forest ecosystem, and contains high levels of biodiversity (N'Goran et al. 2010). The park is a low elevation tropical humid rainforest. Elevation in the southeastern area is approximately 100m with gently rolling hills while in the north, the elevation is approximately 400m in the north with steep ridges (Peal & Kranz 1990). In addition to the western chimpanzee, other endangered and vulnerable species inhabit the site, including forest elephants (Loxodonta africana), pygmy hippopotamus (Hexaprotodon liberiensis), Jentink’s duiker (Cephalophus jentinki), red colobus (Piliocolobus badius), and Diana monkeys (Cercopithecus diana diana, N'Goran 2010).

Table 1. Basic site information for Sapo National Park

Species 'Pan troglodytes verus
Area 1,804 km²
Coordinates Lat: 5.378432 , Lon: -8.496117
Type of site Protected area (National Park)
Habitat types Subtropical/tropical moist lowland
Type of governance

IUCN habitat categories Site designations

Ape status

A survey in 1982 (one year before the establishment of the park), confirmed the presence of chimpanzees in the Sapo forest (Anderson et al. 1983). Based on the estimates from two surveys, one in 2009 (N'Goran et al. 2010) and a second one in 2017 (Tweh et al. 2018), the chimpanzee population in the park has remained relatively stable, with an estimated abundance of approximately 1,055 individuals.

Table 2. Ape population estimates reported for Sapo National Park

Species Year Occurrence Encounter or vistation rate (nests/km; ind/day) Density estimate [ind./ km²] (95% CI) Abundance estimate (95% CI) Survey area Sampling method Analytical framework Source Comments A.P.E.S. database ID
Pan troglodytes verus 1983 Present 0.24 Southeastern sector of the park (50 km²) Line transects Anderson et al. 1983 Total survey effort: 42.7 km
Pan troglodytes verus 2002 Present Sapo National Park Line transects Waitkuwait 2003 Assessment of Fauna & Flora International's bio-monitoring programme
Pan troglodytes verus 2007-2009 0.27 Sapo National Park, excluding south east area Line transects Vogt 2011 Fauna & Flora International bio-monitoring programme
Pan troglodytes verus 2009 4.05 0.86 1079 (CI: 713-1633) Sapo National Park, excluding mining areas Line transects N'Goran et al. 2010
Pan troglodytes verus 2016-2017 0.83 1055 (CI: 595-1870) Sapo National Park, excluding south east area Line transects Tweh et al. 2018 Total survey effort: 38.38 km


Threats

Sapo National Park has been primarily threatened by illegal hunting and mining (Tweh et al. 2018; Greengrass 2015; N'Goran et al. 2010). An estimated 18,000 illegal miners were inhabiting the park in 2010, the majority of which was evicted by the government on the same year (Vogt 2011). A survey of two commercial hunting camps bordering the park revealed high hunting pressure in the area, and the majority of bushmeat harvested was destined to urban areas (Greengrass 2015). The carcasses documented during this survey included chimpanzees as well as other endangered and vulnerable species, such as the red colobus monkey, Diana monkey, and pygmy hippopotamus. Furthermore, the development of the road network around the park is expected to increase hunting pressure and facilitate the bushmeat trade (Greengrass 2015), as well as other illegal activities in the park.

Table 3. Threats to apes reported for Sapo National Park

Category Specific threats Threat level Description Year of threat
10 Geological events Absent
12 Other threat Absent
1 Residential & commercial development 1.1 Residential areas High (more than 70% of population affected) Illegal settlement of miners (Vogt 2011). 2010
3 Energy production & mining 3.2.3 Artisanal mining High (more than 70% of population affected) Illicit gold mining which has decreased since 2010 (Tweh et al. 2018); artisanal mining is still present (Junket per. comm. 2019). Ongoing ( 2019)
5 Biological resource use 5.1 Hunting & collecting terrestrial animals High (more than 70% of population affected) Poaching represents a major threat to chimpanzees and other species in the park (Tweh et al. 2018, N’Goran et al. 2010, Greengrass 2015), and most of the bushmeat is destined to urban areas (Greengrass 2015). Ongoing (2010, 2015, 2018)
6 Human intrusions & disturbance 6.2 War, civil unrest & military exercises High (more than 70% of population affected) Two civil wars since the establishment of the park disrupted conservation activities, and led to illegal occupation of the park, as well as poaching and extraction of natural resources (Greengrass 2015, Collen et al. 2011). 1989-1996, 1999-2003
2 Agriculture & aquaculture 2.1 Annual & perennial non-timber crops Present (unknown severity) Present as a result of illegal settlements; scale unknown (N’Goran et al. 2010). Ongoing (2010)
4 Transportation & service corridors 4.1 Roads & railroads Present (unknown severity) Development of the road network around the park facilitates illegal human activities in the park (Greengrass 2015). Ongoing (2015)
7 Natural system modifications Unknown
8 Invasive & other problematic species, genes & diseases Unknown
9 Pollution Unknown
11 Climate change & severe weather Unknown

IUCN Threats list

  • Illegal mining activity in Sapo © Ben Tally
  • Mining camp © Ben Tally
  • Illegal mining © Ben Tally
  • Illegal mining © Ben Tally

Conservation activities

The Forestry Development Authority of Liberia is responsible for the sustainable management of the forest sector and the protection of all natural resources. It runs the Sapo National Park in collaboration with Fauna & Flora International (FFI) and Wild Chimpanzee Foundation (WCF). Main activities at the national park level include anti-poaching, conservation education awareness, and bio-monitoring & scientific research. As of 2019, the main activities in the research area have been camera trappings (2019,2020, 2021) by FFI and FDA in the entire national park. eDNA (focusing on Pygmy hippopotamus) was conducted in 2022. The national park forms part of the Tai-Grebo-Sapo Forest Complex, which is a conservation priority in West Africa. Conservation efforts in Sapo National Park have mainly focused on law enforcement, conservation awareness, and bio-monitoring. The WCF has supported Community Watch Teams (CWT), which comprise members from surrounding communities, and regularly patrol and support FDA rangers (WCF 2019). CWTs have played an important role in the eviction of illegal miners from the national park (WCF 2019). In 2012, Fauna & Flora International established a long-term bio-monitoring program to follow the population trends for chimpanzees, pygmy hippopotamuses, elephants, as well as duikers, birds, reptiles, and amphibians (Tweh et al. 2018). Together with Liberia's Forestry Development Authority, permanent transects are surveyed twice a year as part of this long-term bio-monitoring program (Tweh et al. 2018).

Table 4. Conservation activities reported for Sapo National Park

Category Specific activity Description Implementing organization(s) Year of activity
2 Counter-wildlife crime 2.3 Conduct regular anti-poaching patrols Community Watch Teams supported by the WCF regularly patrol the site (WCF 2019) Ongoing (2019)
2 Counter-wildlife crime 2.8 Provide training to anti-poaching ranger patrols Members of the Community Watch Teams are trained in the use of equipment (GPS, compass, camera) and patrolling (WCF 2019) Ongoing (2019)
2 Counter-wildlife crime 2.11 Implement monitoring surveillance strategies (e.g., SMART) or use monitoring data to improve effectiveness of patrols Long-term bio-monitoring of chimpanzee population (Tweh et al. 2018) Ongoing (2019)
4 Education & awareness 4.2 Involve local community in ape research and conservation management As part of a long-term bio-monitoring program, staff of Liberia’s Forestry Development Authority and members of the local community are involved in the surveys (Tweh et al. 2018) Ongoing (2018)
5 Protection & restoration 5.9 Resettle illegal human communities (i.e., in a protected area) to another location Eviction of 18,000 illegal settlers in the park by the Liberia's government (Vogt 2011) 2010
5 Protection & restoration 5.2 Legally protect ape habitat The area is designated as National Park (N’Goran et al. 2010) Ongoing (2010)

Conservation activities list (Junker et al. 2017)

Challenges

The influx of local community dwellers inside the national park is hampering the SNP management to implement the protection of chimpanzees. Low manpower for conducting anti-poaching patrols in and outside the park has put chimpanzees under serious threat.

Table 5. Challenges reported for Sapo National Park

Challenges Specific challenges Source Year(s)
4 Institutional support 4.1 Lack of law enforcement Greengrass 2015, N'Goran et al. 2010
2 Resources and capacity 2.5 Lack of equipment/transportation Tweh et al. 2018
1 Site management 1.2 Need for improved coordination Waitkuwait 2003
1 Site management 1.4 Conflict on land tenure Tally, B. pers. comm. 2022


Enablers

Table 6. Enablers reported for Sapo National Park

Enablers Specific enablers Source Year(s)
1 Site management
2 Resources and capacity
3 Engaged community
4 Institutional support
5 Ecological context
6 Safety and stability


Research activities

Several surveys have been done in the park to monitor the chimpanzee population (e.g., N'Goran et al. 2010, Tweh et al. 2018), assess the impacts of conservation interventions (Tweh et al. 2018), investigate the behavior and ecology of chimpanzees in the park (Anderson et al. 1983), and investigate the impact of hunting pressure in the area (Greengrass 2015).


Documented behaviours

Table 7. Behaviours documented for Sapo National Park

Behavior Source
Nut cracking Anderson et al. 1983


Exposure to climate change impacts

External links

Relevant datasets

References

Tweh, C., Kouakou, C.Y., Chira, R., Freeman, B., Githaiga, J.M., Kerwillain, S., Molokwu-Odozi, M., Varney M. and Junker, J. 2018. Nest counts reveal a stable chimpanzee population in Sapo National Park, Liberia. Primate Conservation 2018 (32): 12 pp.

N’Goran, K. P., Kouakou, C.Y. and Herbinger I. 2010. Report on the Population Survey and Monitoring of Chimpanzee in Sapo National Park, Liberia (June–December 2009). Report. Wild Chimpanzee Foundation, Abidjan, Côted’Ivoire.

Anderson, R., Williamson, E.A., and Carter, J. 1983. Chimpanzees of Sapo Forest, Liberia: density, nests, tools and meat-eating. PRIMAaXS, 24(4): 594-601.

Waitkuwait, W.E. 2003. Report on the First Year of Operation of a Community-based Bio-monitoring Programme in and around Sapo National Park, Sinoe County, Liberia. Report. Fauna and Flora International.

Vogt, M. 2011. Results of Sapo National Park Bio-Monitoring Programme 2007-2009. Report. Fauna & Flora International, Monrovia, Liberia.

Greengrass, E. 2015. Commercial hunting to supply urban markets threatens mammalian biodiversity in Sapo National Park. Oryx 50(3), 397–404.

Collen, B., Howard, B., Konie, J., Daniel, O., and Rist, J. 2011. Field surveys for the endangered pygmy hippopotamus Choerpsis liberiensis in Sapo National Park, Liberia. Oryx, 45(1), 35–37.

Wild Chimpanzee Foundation. 2019. Activities of the Wild Chimpanzee Foundation for improved conservation of chimpanzees and their habitat in West Africa. Annual Report 2018.

Peal, A. L., & Kranz, K. R. (1990). Antelopes: GLobal Survey and Regional Action Plans, Part 3. West and

Central Africa. Gland, Switzerland: World Conservation Union.


Page created by: Ben Tally & A.P.E.S. Wiki Team Date: NA