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- Text paragraphs and images are not restricted, so text format and images can be chosen as desired.  
 
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     rather than several keywords such as "Industrial mining", "Large-scale mining", and "Mining".  
 
     rather than several keywords such as "Industrial mining", "Large-scale mining", and "Mining".  
 
-->
 
-->
 
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<!-- Please add information for this site -->
 
[[West Africa]] > [[Republic of Guinea]] > [[Haut Niger National Park]]
 
[[West Africa]] > [[Republic of Guinea]] > [[Haut Niger National Park]]
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SITE UNDER PROGRESS
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= Summary =  <!-- An overview of the site, with a one sentence overview of each of the following sections. can include a site map -->
 
= Summary =  <!-- An overview of the site, with a one sentence overview of each of the following sections. can include a site map -->
   −
* Western chimpanzees ([https://www.iucnredlist.org/species/15935/17989872 Pan troglodytes verus]) are present in the Haut Niger National Park.
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* Western chimpanzees ([https://www.iucnredlist.org/species/15935/17989872 ''Pan troglodytes verus'']) are present in Haut Niger National Park.  
 
* It has been estimated that more than 400 chimpanzees occur in the park.  
 
* It has been estimated that more than 400 chimpanzees occur in the park.  
* The chimpanzee population trend is not known, but there is evidence that it is decreasing.
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* The chimpanzee population trend is not known, but there is evidence that it is decreasing.  
 
* This site has a total size of 12,000 km².  
 
* This site has a total size of 12,000 km².  
* Key threats to chimpanzees are hunting, agriculture and presence of humans in the core area for fishing and honey collection.
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* Key threats to chimpanzees are hunting, agriculture and presence of humans in the core area for fishing and honey collection.  
* The Chimpanzee Conservation Center rehabilitates orphaned chimpanzees and released a group into the wild in 2008. It also conducts an awareness raising program.  
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* The Chimpanzee Conservation Center rehabilitates orphaned chimpanzees and released a group into the wild in 2008. It also conducts an awareness raising program.
 
* The site protects one of the last open dry forests in West Africa and serves key hydrological functions. It features a high biodiversity including lions, hippos and a diversity of fish and bird species, and contains a UNESCO Biosphere Reserve, an Important Bird Area and two Ramsar sites.
 
* The site protects one of the last open dry forests in West Africa and serves key hydrological functions. It features a high biodiversity including lions, hippos and a diversity of fish and bird species, and contains a UNESCO Biosphere Reserve, an Important Bird Area and two Ramsar sites.
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[[Image:Niger_River.jpg|right|thumb|Niger River]]
      
= Site characteristics = <!-- A paragraph summary of physical and geographic aspects of the site, and a table of key information -->
 
= Site characteristics = <!-- A paragraph summary of physical and geographic aspects of the site, and a table of key information -->
    
The Haut Niger National Park (also called Upper Niger National Park) is situated in the Niger basin in east-central Guinea. It has been designated a National Park in 1997 and consists of two distinct core areas, Mafou forest (554 km²) and Kouyah forest (674 km²), and a buffer zone (Fleury-Brugière & Brugière 2010). The total area of the park, meaning core areas and buffer zone, is around 12,000 km2 (Fleury-Brugière & Brugière 2010). The two core areas are designated as IUCN category II and the buffer zone is designated as IUCN category V (Brugière & Kormos 2009).  
 
The Haut Niger National Park (also called Upper Niger National Park) is situated in the Niger basin in east-central Guinea. It has been designated a National Park in 1997 and consists of two distinct core areas, Mafou forest (554 km²) and Kouyah forest (674 km²), and a buffer zone (Fleury-Brugière & Brugière 2010). The total area of the park, meaning core areas and buffer zone, is around 12,000 km2 (Fleury-Brugière & Brugière 2010). The two core areas are designated as IUCN category II and the buffer zone is designated as IUCN category V (Brugière & Kormos 2009).  
   
In 2002 Mafou forest and surrounding areas were proclaimed a UNESCO Biosphere Reserve ([http://www.unesco.org/new/en/natural-sciences/environment/ecological-sciences/biosphere-reserves/africa/guinea/haut-niger/  UNESCO MAB 2011]) consisting of the Mafou forest as the core area, a buffer zone (3,641 km2) and transition area (2,275 km2, Brugière & Kormos 2009).  
 
In 2002 Mafou forest and surrounding areas were proclaimed a UNESCO Biosphere Reserve ([http://www.unesco.org/new/en/natural-sciences/environment/ecological-sciences/biosphere-reserves/africa/guinea/haut-niger/  UNESCO MAB 2011]) consisting of the Mafou forest as the core area, a buffer zone (3,641 km2) and transition area (2,275 km2, Brugière & Kormos 2009).  
   
Haut Niger is located at the border of the two prefectures Faranah and Kankan, on a plateau with stream valleys. The annual rainfall ranges from 1,500 and 1,700 mm (BirdLife International 2018). This National Parks protects one of the last primary open dry forests in West Africa (UNESCO MAB 2011). The habitat is characterized by wooded and open savanna, dry forest, bamboo forest, and gallery forest along the Niger River and its tributaries (UNESCO MAB 2011, Fleury-Brugière & Brugière 2010). There are also permanent ponds which are an important part of cultural life (Ramsar 2002). Haut Niger National Park is characterized by a high biodiversity. With its 300 bird species (Nikolaus 1999) Mafou forest has been declared an Important Bird Area ([http://datazone.birdlife.org/site/factsheet/mafou-iba-guinea/text  BirdLife International 2018]).  
 
Haut Niger is located at the border of the two prefectures Faranah and Kankan, on a plateau with stream valleys. The annual rainfall ranges from 1,500 and 1,700 mm (BirdLife International 2018). This National Parks protects one of the last primary open dry forests in West Africa (UNESCO MAB 2011). The habitat is characterized by wooded and open savanna, dry forest, bamboo forest, and gallery forest along the Niger River and its tributaries (UNESCO MAB 2011, Fleury-Brugière & Brugière 2010). There are also permanent ponds which are an important part of cultural life (Ramsar 2002). Haut Niger National Park is characterized by a high biodiversity. With its 300 bird species (Nikolaus 1999) Mafou forest has been declared an Important Bird Area ([http://datazone.birdlife.org/site/factsheet/mafou-iba-guinea/text  BirdLife International 2018]).  
   
Two Ramsar sites were gazetted that same year. The Niger-Mafou Ramsar site has a  total area of 10,154.50 km2 and is characterized by permanent and seasonal rivers, and marshes ([https://rsis.ramsar.org/ris/1163  Ramsar 2002]). Migratory large mammals use this area as a corridor with abundant water sources year round and the site is visited by birds (Ramsar 2002). The second site, Niger-Niandan-Milo covers an area of 10,464 km2 and features a high diversity in fish species (Crespi 1998) and serves important hydrological functions ([https://rsis.ramsar.org/ris/1164  Ramsar 2017]). The site is also known for its diversity in waterbirds (Ramsar 2017).  
 
Two Ramsar sites were gazetted that same year. The Niger-Mafou Ramsar site has a  total area of 10,154.50 km2 and is characterized by permanent and seasonal rivers, and marshes ([https://rsis.ramsar.org/ris/1163  Ramsar 2002]). Migratory large mammals use this area as a corridor with abundant water sources year round and the site is visited by birds (Ramsar 2002). The second site, Niger-Niandan-Milo covers an area of 10,464 km2 and features a high diversity in fish species (Crespi 1998) and serves important hydrological functions ([https://rsis.ramsar.org/ris/1164  Ramsar 2017]). The site is also known for its diversity in waterbirds (Ramsar 2017).  
 
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In a survey from 1995 to 1997 a total of 94 species of mammals were identified (Ziegler et al. 2002). The park is home to a small population of lions (''Panthera leo'') ([http://lionalert.org/alert/lions_in/Guinea  Lion Alert 2012], Bauer 2003), as well as leopards (''Panthera pardus''), hippopotamus (''Hippopotamus amphibius'', Brugière et al. 2006), jackals (''Canis sp.''), civets, patas monkeys (''Erythrocebus patas''), baboons (''Papio sp.''), green monkeys (''Chlorocebus sabaeus''), and mangabeys (''Cercocebus sp.'', Brugière  2006, Brugière & Kormos 2009, WCF 2012, Ziegler et al. 2002).
In a survey from 1995 to 1997 a total of 94 species of mammals were identified (Ziegler et al. 2002). The park  
  −
is home to a small population of lions (Panthera leo) ([http://lionalert.org/alert/lions_in/Guinea  Lion Alert 2012], Bauer 2003), as well as leopards, hippos (Brugière et al. 2006), jackals, civets, patas monkeys, baboons, green monkeys, and mangabeys (Brugière  2006, Brugière & Kormos 2009, WCF 2012, Ziegler et al. 2002).
         
'''Table 1: Basic site information for Haut Niger National Park'''
 
'''Table 1: Basic site information for Haut Niger National Park'''
 
{| border="1" cellpadding="5" cellspacing="0" class="basic-information"
 
{| border="1" cellpadding="5" cellspacing="0" class="basic-information"
| Area:             <!-- Please include units: km2/ha e.g 200ha    -->
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|Area            <!-- Please include units: km2/ha e.g 200ha    -->
|12,000 km²  
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|12,000 km²
 
|-
 
|-
| Designation:     <!-- Protected area, Forestry concession, Community reserve   -->
+
|Coordinates
 +
|10.21 N, -10.09 W
 +
|-
 +
|Designation      <!-- National Park, Nature Reserve, etc.   -->
 
|National Park
 
|National Park
 
|-
 
|-
|Habitat types:   <!-- List IUCN Habitat Classification 3.0 categories present (Without number), see link below -->
+
|Habitat types    <!-- List IUCN Habitat Classification 3.0 categories present (Without number), see link below -->
|Subtropical/tropical dry forest, dry savanna, subtropical/tropical dry shrubland, permanent rivers, permanent freshwater marshes/pools  
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|Subtropical/tropical dry forest, dry savanna, subtropical/tropical dry shrubland, permanent rivers, permanent freshwater marshes/pools, arable land
 
|}
 
|}
 
[http://www.unitar.org/hiroshima/sites/unitar.org.hiroshima/files/Annex%201%20-%20IUCN%20Classification%20Schemes.pdf IUCN habitat categories]  [[Site designations]]
 
[http://www.unitar.org/hiroshima/sites/unitar.org.hiroshima/files/Annex%201%20-%20IUCN%20Classification%20Schemes.pdf IUCN habitat categories]  [[Site designations]]
 
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<br>
    
= Ape status =  <!-- a text overview of ape status (population sizes, trends etc), followed by a table of specific surveys and results -->
 
= Ape status =  <!-- a text overview of ape status (population sizes, trends etc), followed by a table of specific surveys and results -->
   −
In a nationwide survey from 1995 to 1997 Rebecca Kormos identified Mafou forest as an “area of particular interest thought to have a healthy population of chimpanzees” based on reconnaissance surveys (Ham 1998). From 2001 to 2002 transect surveys were conducted in the Mafou core area and a total abundance of 480 individuals was estimated for that core area (Fleury-Brugière & Brugière 2010). Extrapolated to the entire park it has been suggested that the park protects up to 650 chimpanzees (Kormos et al. 2003). A follow-up survey conducted between 2008 and 2009 in the northern part of Mafou found a lower chimpanzee density and derived an estimate of 334 individuals (200-601) for Mafou (Humle et al. 2009a). It was concluded that the population likely decreased (Humle et al. 2009a). Yet another transect survey in 2009 found even lower densities for Mafou and estimated 288 individuals ([http://www.wildchimps.org  WCF 2012]). At the same time Kouyah core area was sampled for the first time and 132 individuals were estimated to occur there (WCF 2012).
+
In a nationwide survey from 1995 to 1997 Rebecca Kormos identified Mafou forest as an “area of particular interest thought to have a healthy population of chimpanzees” based on reconnaissance surveys (Ham 1998). From 2001 to 2002 transect surveys were conducted in the Mafou core area and a total abundance of 480 individuals was estimated for that core area (Fleury-Brugière & Brugière 2010). Extrapolated to the entire park it has been suggested that the park protects up to 650 chimpanzees (Kormos et al. 2003). A follow-up survey conducted between 2008 and 2009 in the northern part of Mafou found a lower chimpanzee density and derived an estimate of 334 individuals (200-601) for Mafou (Humle et al. 2009a). It was concluded that the population likely decreased (Humle et al. 2009a). Yet another transect survey in 2009 found even lower densities for Mafou and estimated 288 individuals (190-437) ([http://www.wildchimps.org  WCF 2012]). At the same time Kouyah core area was sampled for the first time and 132 individuals (60-290) were estimated to occur there (WCF 2012).
 
   
Haut Niger National Park was identified as an ‘Exceptionally Important Priority Area’ in the Regional Conservation Action Plan for western chimpanzees in 2003 (Kormos & Boesch 2003).
 
Haut Niger National Park was identified as an ‘Exceptionally Important Priority Area’ in the Regional Conservation Action Plan for western chimpanzees in 2003 (Kormos & Boesch 2003).
   Line 68: Line 59:  
! Species
 
! Species
 
! Year
 
! Year
! Abundance estimate (95% Confidence Intervall)
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! Abundance estimate (95% confidence interval)
! Density estimate (per km2)
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! Density estimate (per km²)
! Encounter rate
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! Encounter rate (nests/km)
 
! Area
 
! Area
 
! Method
 
! Method
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! A.P.E.S. database ID
 
! A.P.E.S. database ID
 
|-
 
|-
|Western chimpanzee
+
|''Pan troglodytes verus''
 
|1995-1997
 
|1995-1997
|
+
|XXXX (abundance est.)
|
+
|XXXX (density est.)
|presence
+
|Present
 
|Mafou forest
 
|Mafou forest
|reconnaissance survey
+
|Index survey
 
|Ham 1998
 
|Ham 1998
|
+
|Reconnaissance survey
 
|
 
|
 
|-
 
|-
|Western chimpanzee
+
|''Pan troglodytes verus''
 
|1995-1997
 
|1995-1997
|
+
|XXXX (abundance est.)
|
+
|XXXX (density est.)
|presence
+
|Present
 
|Kouyah forest
 
|Kouyah forest
|anectodal reports
+
|Informed guess
 
|Ham 1998
 
|Ham 1998
|
+
|Anectodal reports
 
|
 
|
 
|-
 
|-
|Western chimpanzee
+
|''Pan troglodytes verus''
 
|2001-2002
 
|2001-2002
 
|480 weaned individuals
 
|480 weaned individuals
 
|0.87 weaned individuals/km²
 
|0.87 weaned individuals/km²
|7.9 nests/km (range: 0.16–11.7)
+
|7.9 (range: 0.16–11.7)
 
|Mafou core area
 
|Mafou core area
|transect survey, DISTANCE analysis
+
|Line transects (Distance)
 
|Fleury-Brugière & Brugière 2010
 
|Fleury-Brugière & Brugière 2010
|total survey effort: 103.83 km
+
|Total survey effort: 103.83 km
 
|
 
|
 
|-
 
|-
|Western chimpanzee
+
|''Pan troglodytes verus''
 
|2008-2009
 
|2008-2009
|334 individuals (CI: 200-601)
+
|334 (200-601)
|0.60 individuals/km2 (CI: 0.36-1.1)
+
|0.60 (0.36-1.1)
|7.6 nests/km
+
|7.6
 
|Mafou core area
 
|Mafou core area
|transect survey, DISTANCE analysis
+
|Line transects (Distance)
|Humle et al. 2009
+
|Humle et al. 2009a
|total survey effort: 60.5 km
+
|Total survey effort: 60.5 km
 
|
 
|
 
|-
 
|-
|Western chimpanzee
+
|''Pan troglodytes verus''
 
|2009
 
|2009
|Mafou: 288 individuals (CI: 190-437)
+
|Mafou: 288 (190-437), Kouyah: 132 (60-290)
 
+
|Mafou: 0.47, Kouyah: 0.23
Kouyah: 132 individuals (CI: 60-290)
+
|2.34
 
+
|Mafou + Kouya core area
|Mafou: 0.47 individuals/km2
+
|Line transects (Distance)
 
  −
Kouyah: 0.23 individuals/km2
  −
|2.34 nests/km
  −
|Mafou + Kouyah core area
  −
|transect survey, DISTANCE analysis
   
|WCF 2012
 
|WCF 2012
|total survey effort: 196 km
+
|Total survey effort: 196 km
 
|
 
|
 
|}
 
|}
      
= Threats =    <!-- a text overview of threats, followed by a table of key threats -->
 
= Threats =    <!-- a text overview of threats, followed by a table of key threats -->
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The main threats to chimpanzees in Haut Niger National Park are hunting, agriculture and presence of humans in the core area for fishing and honey collection. Humle et al. (2009) identified hunting as one of the most prevalent pressures in the park with an encounter rate of poaching signs of 0.15/km. A similar encounter rate was found by the WCF survey with 0.18 signs/km (WCF 2012). Chimpanzees are also hunted opportunistically. In the bushmeat survey in 2001 Brugière and Magassouba (2009) found 14 chimpanzee carcasses offered for sale over the course of a year.
 
The main threats to chimpanzees in Haut Niger National Park are hunting, agriculture and presence of humans in the core area for fishing and honey collection. Humle et al. (2009) identified hunting as one of the most prevalent pressures in the park with an encounter rate of poaching signs of 0.15/km. A similar encounter rate was found by the WCF survey with 0.18 signs/km (WCF 2012). Chimpanzees are also hunted opportunistically. In the bushmeat survey in 2001 Brugière and Magassouba (2009) found 14 chimpanzee carcasses offered for sale over the course of a year.
 
Hunters regularly set fires at night as a hunting technique to drive out animals so they can be shot while fleeing. 70% of transects surveyed by Humle et al. (2009) showed signs of recent burning.  
 
Hunters regularly set fires at night as a hunting technique to drive out animals so they can be shot while fleeing. 70% of transects surveyed by Humle et al. (2009) showed signs of recent burning.  
   
Slash and burn agriculture is another threat to chimpanzees (Humle et al. 2009a). According to the WCF survey (2012) agriculture is widespread in the Kouyah core area, but can only be found in the south of Mafou core area in proximity to two villages. The WCF (2012) encountered 0.31 signs of agriculture per km across the two parks. With around 80,000 people living in the buffer zone of the park, agriculture is widespread there and has led to severe environmental degradation (Ramsar 2017).  
 
Slash and burn agriculture is another threat to chimpanzees (Humle et al. 2009a). According to the WCF survey (2012) agriculture is widespread in the Kouyah core area, but can only be found in the south of Mafou core area in proximity to two villages. The WCF (2012) encountered 0.31 signs of agriculture per km across the two parks. With around 80,000 people living in the buffer zone of the park, agriculture is widespread there and has led to severe environmental degradation (Ramsar 2017).  
   
People regularly enter the core areas to fish and collect honey. For the latter, trees are felled for the harvest of wild honey and self-made bee hives are set up in trees in the periphery of Mafou core area (Humle et al. 2009a). Fishing is still partly done using techniques destructive for waterways and fish species, such as using explosives and chemicals, leading to high fish mortalities (Ramsar 2017).  
 
People regularly enter the core areas to fish and collect honey. For the latter, trees are felled for the harvest of wild honey and self-made bee hives are set up in trees in the periphery of Mafou core area (Humle et al. 2009a). Fishing is still partly done using techniques destructive for waterways and fish species, such as using explosives and chemicals, leading to high fish mortalities (Ramsar 2017).  
   
While there a no permanent human settlements within the two core areas, fishing camps can be found along the rivers and they harbor up to 100 people for up to six months (Humle et al. 2009a, WCF 2012).  
 
While there a no permanent human settlements within the two core areas, fishing camps can be found along the rivers and they harbor up to 100 people for up to six months (Humle et al. 2009a, WCF 2012).  
   
Illegal logging was very prevalent in the buffer zone until 2009, but due to increased law enforcement activities, including road blocks, such activities ceased (Humle et al. 2009a). But these initiatives were short-term, so the current impact of this threat is unclear.  
 
Illegal logging was very prevalent in the buffer zone until 2009, but due to increased law enforcement activities, including road blocks, such activities ceased (Humle et al. 2009a). But these initiatives were short-term, so the current impact of this threat is unclear.  
   
In conclusion though, after a nationwide survey of 11 sites a WCF report concluded that anthropogenic signs inside the two core areas were a lot less prevalent than in other protected areas across Guinea (WCF 2012).  
 
In conclusion though, after a nationwide survey of 11 sites a WCF report concluded that anthropogenic signs inside the two core areas were a lot less prevalent than in other protected areas across Guinea (WCF 2012).  
      −
<br>
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'''Table 3: Threats to great apes in Haut Niger National Park'''
'''Table 3: Threats to great apes in SITE NAME'''
   
{| border="1" cellpadding="5" cellspacing="0" class="threats-table"
 
{| border="1" cellpadding="5" cellspacing="0" class="threats-table"
 
!align="left"|Category  <!-- Do not change threat categories -->
 
!align="left"|Category  <!-- Do not change threat categories -->
 
!Specific threats    <!-- For specific threats, please use list of comma separated keywords from the list linked below -->
 
!Specific threats    <!-- For specific threats, please use list of comma separated keywords from the list linked below -->
 
!Threat level        <!-- For threat level, please use keywords: unknown, low, high -->
 
!Threat level        <!-- For threat level, please use keywords: unknown, low, high -->
 +
!Quantified severity <!-- e.g., encounter rate, number of miners etc. (with reference)-->
 
!Description        <!-- You can add descriptive information here -->
 
!Description        <!-- You can add descriptive information here -->
!Year of threat
+
!Year of threat     <!-- if ongoing or unknown add year of reference in brackets-->
 
|-
 
|-
 
|1. Residential & commercial development
 
|1. Residential & commercial development
 
|1.1. Housing & urban areas
 
|1.1. Housing & urban areas
|medium
+
|Medium
|fishing camps along the rivers, also inside core areas, with up to 100 people living in these camps for up to 6 months [11, 23]
+
|0.01 villages/km (1 village, survey effort: 195.691 km, WCF 2012)
|on-going
+
|Fishing camps along the rivers, also inside core areas, with up to 100 people living in these camps for up to 6 months (Humle et al. 2009a, WCF 2012)
 +
|Ongoing (2009, 2012)
 
|-
 
|-
 
|2. Agriculture & aquaculture
 
|2. Agriculture & aquaculture
 
|2.1. Annual & Perennial Non-Timber Crops
 
|2.1. Annual & Perennial Non-Timber Crops
|medium
+
|Medium
|signs of agricultural throughout Kouyah core area [23] and widespread in buffer zone [11, 23]; people also use trees in the periphery of Mafou core area for setting up self-made bee hives [11]
+
|0.31 signs of agriculture/km (60 signs of agriculture, survey effort: 195.691 km, WCF 2012)
|on-going
+
|Signs of agricultural throughout Kouyah core area (WCF 2012) and widespread in buffer zone (Humle et al. 2009a, WCF 2012); people also use trees in the periphery of Mafou core area for setting up self-made bee hives (Humle et al. 2009a)
 +
|Ongoing (2009, 2012)
 
|-
 
|-
 
|3. Energy production & mining
 
|3. Energy production & mining
 
|
 
|
|absent
+
|Absent
 +
|
 
|
 
|
 
|
 
|
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|4. Transportation & service corridors
 
|4. Transportation & service corridors
 
|4.1. Roads & railroads
 
|4.1. Roads & railroads
|medium
+
|Medium
|trails and path can be found also in the core area and these are used by hunters and fishers [11, 23], the threat from such trails to chimpanzees is indirect as they facilitate access for humans
+
|0.26 roads/km (51 roads, survey effort: 195.691 km, WCF 2012)
|on-going
+
|Trails and path can be found also in the core area and these are used by hunters and fishers (Humle et al 2009a, WCF 2012), the threat from such trails to chimpanzees is indirect as they facilitate access for humans
 +
|Ongoing (2009, 2012)
 
|-
 
|-
 
|5. Biological resource use
 
|5. Biological resource use
|5.1.Hunting & collecting terrestrial animals
+
|5.1. Hunting & collecting terrestrial animals
|high
+
|High
|widespread hunting especially in the buffer zone but also in the core areas [11, 23], chimpanzees also hunted opportunistically [6]; wild honey collection [11]
+
|0.18 signs of hunting/km (36 signs of hunting, survey effort: 195.691 km, WCF 2012)
|on-going
+
|Widespread hunting especially in the buffer zone but also in the core areas (Humle et al 2009a, WCF 2012), chimpanzees also hunted opportunistically (Brugière and Magassouba 2009); wild honey collection (Humle et al. 2009a)
 +
|Ongoing (2009, 2012)
 
|-
 
|-
 
|
 
|
 
|5.3. Logging & wood harvesting
 
|5.3. Logging & wood harvesting
|medium
+
|Medium
|illegal logging in the buffer zone [11]
+
|0.03 signs of wood extraction/km (5 signs of wood extraction, survey effort: 195.691 km, WCF 2012)
|likely on-going
+
|Illegal logging in the buffer zone (Humle et al. 2009a)
 +
|Likely ongoing (2009)
 
|-
 
|-
 
|
 
|
 
|5.4. Fishing & harvesting aquatic resources
 
|5.4. Fishing & harvesting aquatic resources
|low
+
|Low
|fishing in the rivers, also using dynamite and chemicals, is widespread [21]; effect on chimpanzees is indirect due to presence of humans in the core area
+
|
|on-going
+
|Fishing in the rivers, also using dynamite and chemicals, is widespread (Ramsar 2017); effect on chimpanzees is indirect due to presence of humans in the core area
 +
|Ongoing (2017)
 
|-
 
|-
|6. Human intrusion & disturbance
+
|6. Human intrusions & disturbance
 +
|
 +
|Absent
 
|
 
|
|absent
   
|
 
|
 
|
 
|
Line 214: Line 202:  
|7. Natural system modifications
 
|7. Natural system modifications
 
|7.1. Fire & Fire suppression
 
|7.1. Fire & Fire suppression
|medium
+
|Medium
|fires are set at night by hunters to shoot fleeing animals [11], can effect behavior of chimpanzees
+
|
|on-going
+
|Fires are set at night by hunters to shoot fleeing animals (Humle et al. 2009a), can effect behavior of chimpanzees
 +
|Ongoing (2009)
 
|-
 
|-
 
|8. Invasive & other problematic species, genes, diseases
 
|8. Invasive & other problematic species, genes, diseases
 
|
 
|
|unknown
+
|Unknown
 +
|
 
|
 
|
 
|
 
|
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|9. Pollution
 
|9. Pollution
 
|
 
|
|absent
+
|Absent
 +
|
 
|
 
|
 
|
 
|
Line 232: Line 223:  
|10. Geological Events
 
|10. Geological Events
 
|
 
|
|absent
+
|Absent
 +
|
 
|
 
|
 
|
 
|
Line 238: Line 230:  
| 11. Climate change & severe weather
 
| 11. Climate change & severe weather
 
|
 
|
|unknown
+
|Unknown
 +
|
 
|
 
|
 
|
 
|
Line 244: Line 237:  
|12. Other options
 
|12. Other options
 
|
 
|
|absent
+
|Absent
 +
|
 
|
 
|
 
|
 
|
 
|-
 
|-
 
|}
 
|}
[[Threats list]]
+
[https://www.iucnredlist.org/resources/threat-classification-scheme IUCN Threats list]
 
      
= Conservation activities =      <!-- a text overview of conservation activities, followed by a table of key activities -->
 
= Conservation activities =      <!-- a text overview of conservation activities, followed by a table of key activities -->
Line 256: Line 249:  
As part of the “Programme Regional d’Aménagement des Bassins Versants du Niger et de la Gambie” (PRABV) which was funded by the European Union in the 1990’s, biodiversity surveys were conducted in and around Mafou and Kouya forest, which ultimately led to the creation of the Haut Niger National Park in 1997 (Brugière & Kormos 2009). From 1995 to 2005 the European funded AGIR project (Programme Régional d’Appui à la Gestion Intégrée des Ressources Naturelles des Bassins du Niger et de la Gambie) implemented a management plan in collaboration with government officials and supported law enforcement within the park (Humle et al. 2009a). When funding ceased government authorities were lacking the financial, logistical and technical means to continue these efforts at the same level (Humle et al. 2009a).  
 
As part of the “Programme Regional d’Aménagement des Bassins Versants du Niger et de la Gambie” (PRABV) which was funded by the European Union in the 1990’s, biodiversity surveys were conducted in and around Mafou and Kouya forest, which ultimately led to the creation of the Haut Niger National Park in 1997 (Brugière & Kormos 2009). From 1995 to 2005 the European funded AGIR project (Programme Régional d’Appui à la Gestion Intégrée des Ressources Naturelles des Bassins du Niger et de la Gambie) implemented a management plan in collaboration with government officials and supported law enforcement within the park (Humle et al. 2009a). When funding ceased government authorities were lacking the financial, logistical and technical means to continue these efforts at the same level (Humle et al. 2009a).  
 
In 2008 the Guinean government stepped up efforts against illegal logging using a nation-wide information campaign via the radio and setting up road blocks in 2008-2009 at the roads entering the national parc (Humle et al. 2009a).  
 
In 2008 the Guinean government stepped up efforts against illegal logging using a nation-wide information campaign via the radio and setting up road blocks in 2008-2009 at the roads entering the national parc (Humle et al. 2009a).  
   
In 1997 the sanctuary [https://www.projetprimates.com/en/  Chimpanzee Conservation Center (CCC)] was founded. The sanctuary rescues, rehabilitates and cares for orphaned chimpanzees. As it is the only chimpanzee sanctuary in Guinea, it is the contact point for taking care of chimpanzees confiscated by the Guinean government officials.
 
In 1997 the sanctuary [https://www.projetprimates.com/en/  Chimpanzee Conservation Center (CCC)] was founded. The sanctuary rescues, rehabilitates and cares for orphaned chimpanzees. As it is the only chimpanzee sanctuary in Guinea, it is the contact point for taking care of chimpanzees confiscated by the Guinean government officials.
 
In 2008 CCC released the first group of chimpanzees in Mafou core area consisting of 12 individuals with the aim of reinforcing the wild chimpanzee population in the park (Projet Primates 2016, Humle et al. 2011). Two years later 9 individuals were still free-living and two females had offsprings (Humle et al. 2011).
 
In 2008 CCC released the first group of chimpanzees in Mafou core area consisting of 12 individuals with the aim of reinforcing the wild chimpanzee population in the park (Projet Primates 2016, Humle et al. 2011). Two years later 9 individuals were still free-living and two females had offsprings (Humle et al. 2011).
Since 2004 CCC also implements an awareness raising program on the topics of ‘importance of chimpanzee conservation’, activities that are illegal and threatening chimpanzees, and the general importance of protecting the national park by holding village meetings, distributing information material and creating radio programs in local languages for broadcasting across Guinea (Projet Primates 2016).  
+
Since 2004 CCC also implements an awareness raising program on the topics of ‘importance of chimpanzee conservation’, activities that are illegal and threatening chimpanzees, and the general importance of protecting the national park by holding village meetings, distributing information material and creating radio programs in local languages for broadcasting across Guinea (Projet Primates 2016). There is also an ecomuseum and a botanical garden in the national park (UNESCO MAB 2011).
   −
There is also an ecomuseum and a botanical garden in the national park (UNESCO MAB 2011).
     −
 
+
'''Table 4: Conservation activities in Haut Niger National Park'''
 
  −
'''Table 3: Conservation activities in Haut Niger National Park'''
   
{| border="1" cellpadding="5" cellspacing="0" class="conservation-actions-table"
 
{| border="1" cellpadding="5" cellspacing="0" class="conservation-actions-table"
 
!align="left"|Category  <!-- Do not change threat categories -->
 
!align="left"|Category  <!-- Do not change threat categories -->
 
!Specific activity    <!-- For specific threats, please use list of comma separated keywords from the list linked below -->
 
!Specific activity    <!-- For specific threats, please use list of comma separated keywords from the list linked below -->
 
!Description        <!-- You can add descriptive information here -->
 
!Description        <!-- You can add descriptive information here -->
!Year of activity        <!-- You can add descriptive information here -->
+
!Year of activity        <!-- if ongoing or unknown add year of reference in brackets -->
 
|-
 
|-
 
|1. Residential & commercial development
 
|1. Residential & commercial development
|absent
+
|Not reported
|
+
|XXXXX (description)
|
+
|XXXXX (year)
 
|-
 
|-
 
|2. Agriculture & aquaculture
 
|2. Agriculture & aquaculture
|absent
+
|Not reported
|
+
|XXXXX (description)
|
+
|XXXXX (year)
 
|-
 
|-
 
|3. Energy production & mining
 
|3. Energy production & mining
|absent
+
|Not reported
|
+
|XXXXX (description)
|
+
|XXXXX (year)
 
|-
 
|-
 
|4. Transportation & service corridors
 
|4. Transportation & service corridors
|absent
+
|Not reported
|
+
|XXXXX (description)
|
+
|XXXXX (year)
 
|-
 
|-
 
|5. Biological resource use
 
|5. Biological resource use
|5.4. Implement road blocks to inspect cars for illegal primate bushmeat  
+
|5.4. Implement road blocks to inspect cars for illegal primate bushmeat
|the road blocks set up to control illegal logging were used to also control illegal bushmeat [11]
+
|The road blocks set up to control illegal logging were used to also control illegal bushmeat (Humle et al. 2009a)
 
|2008-2009
 
|2008-2009
 
|-
 
|-
 
|
 
|
 
|5.6. Conduct regular anti-poaching patrols
 
|5.6. Conduct regular anti-poaching patrols
|
+
|The AGIR project supported law enforcement within the park (Humle et al. 2009a)
|unknown
+
|Unknown (2009)
 
|-
 
|-
|6. Human intrusion & disturbance
+
|6. Human intrusions & disturbance
|absent
+
|Not reported
|
+
|XXXXX (description)
|
+
|XXXXX (year)
 
|-
 
|-
 
|7. Natural system modifications
 
|7. Natural system modifications
|absent
+
|Not reported
|
+
|XXXXX (description)
|
+
|XXXXX (year)
 
|-
 
|-
 
|8. Invasive & other problematic species, genes, diseases
 
|8. Invasive & other problematic species, genes, diseases
|absent
+
|Not reported
|
+
|XXXXX (description)
|
+
|XXXXX (year)
 
|-
 
|-
 
|9. Pollution
 
|9. Pollution
|absent
+
|Not reported
|
+
|XXXXX (description)
|
+
|XXXXX (year)
 
|-
 
|-
 
|10. Education & Awareness  
 
|10. Education & Awareness  
|10.1. Educate local communities about primates and sustainable use  
+
|10.1. Educate local communities about primates and sustainable use
|as part of awareness program by Chimpanzee Conservation Center (CCC) [19]
+
|As part of awareness program by Chimpanzee Conservation Center (CCC, Projet Primates 2016)
|on-going
+
|Ongoing (2016)
 
|-
 
|-
 
|
 
|
|10.4. Regularly play TV & radio announcements to raise primate conservation awareness  
+
|10.4. Regularly play TV & radio announcements to raise primate conservation awareness
|as part of awareness program by CCC [19]
+
|As part of awareness program by CCC (Projet Primates 2016)
|on-going
+
|Ongoing (2016)
 
|-
 
|-
 
|
 
|
|10.5. Implement multimedia campaigns using theatre, film, print media, discussions  
+
|10.5. Implement multimedia campaigns using theatre, film, print media, discussions
|as part of awareness program by CCC [19]
+
|As part of awareness program by CCC (Projet Primates 2016)
|on-going
+
|Ongoing (2016)
 
|-
 
|-
|11. Habitat Protection  
+
|11. Habitat Protection
|11.2. Legally protect primate habitat  
+
|11.2. Legally protect primate habitat
|designated as National Park since 1997 [8]
+
|Designated as National Park since 1997 (Fleury-Brugière and Brugière 2010)
|since 1997
+
|Since 1997
 
|-
 
|-
 
|12. Species Management
 
|12. Species Management
 
|12.15. Reintroduce primates in groups
 
|12.15. Reintroduce primates in groups
|12 individuals released in Mafou core area [11, 19]
+
|12 individuals released in Mafou core area (Humle et al. 2009a, Projet Primates 2016)
 
|2008
 
|2008
 
|-
 
|-
 
|
 
|
|12.24. Rehabilitate injured/orphaned primates  
+
|12.24. Rehabilitate injured/orphaned primates
|CCC takes care of and rehabilitates orphaned chimpanzees (CCC) [19]
+
|CCC takes care of and rehabilitates orphaned chimpanzees (Projet Primates 2016)
|since 1997
+
|Since 1997
 
|-
 
|-
 
|13. Livelihood; Economic & Other Incentives  
 
|13. Livelihood; Economic & Other Incentives  
|13.5. Permanent presence of staff/manager  
+
|13.5. Permanent presence of staff/manager
|permanent presence of CCC staff since 1997 [19]
+
|Permanent presence of CCC staff since 1997 (Projet Primates 2016)
|since 1997
+
|Since 1997
 
|-
 
|-
 
|}
 
|}
[[Conservation activities list]]
+
[[Conservation activities list (Junker et al. 2017)]]
    +
= Impediments = <!-- Overview of impediments to ape conservation -->
    +
Lack of funding as well as logistical and technical support were cited as impediments to making and implementing a new management plan for the park (Humle et al. 2009a). The evaluation of management effectiveness concluded, that due to a lack of financial and technical means as well as lack of training, an efficient monitoring and law enforcement throughout the park was not possible (METT 2009).
   −
= Impediments = <!-- Overview of impediments to ape conservation -->
     −
Lack of funding as well as logistical and technical support were cited as impediments to making and implementing a new management plan for the park (Humle et al. 2009a).
+
'''Table 5: Impediments reported for Haut Niger National Park'''
 +
{| border="1" cellpadding="5" cellspacing="0" class="impediments-table"
 +
!align="left"|Impediment  <!-- Do not change categories -->
 +
!Source  <!-- source for impediment mentioned -->
 +
|-
 +
|Lack of capacity/training
 +
|METT 2009
 +
|-
 +
|Lack of financial means
 +
|Humle et al. 2009a, METT 2009
 +
|-
 +
|Lack of logistical means
 +
|Humle et al. 2009a, METT 2009
 +
|-
 +
|Lack of technical means
 +
|Humle et al. 2009a, METT 2009
 +
|}
 +
 
 +
= Research activities = <!-- Overview of research activities -->
    +
To inform the creation of the National Park several biomonitoring studies have been undertaken targeting different taxa, such as mammals (Ziegler 2002), fish (Crespi 1998), ungulates (Brugière 2005), birds (Nikolaus 1999), and amphibians and reptiles (Greenbaum & Carr 2005).  As detailed above several surveys on chimpanzees have been implemented (Ham 1998, Humle et al. 2009a, Fleury-Brugière & Brugière 2010, WCF 2012).
 +
In addition, studies were conducted that analysed the bushmeat trade in Haut Niger National Park (Brugière & Magassouba 2009, Humle & Alexandre 2015). Following the release of a group of rehabilitated chimpanzees their ranging patterns and behavior has been studies (Humle et al. 2009b, Humle et al. 2011).
   −
= Research activities = <!-- Overview of research activities, and keyword list of documented behaviours -->
     −
To inform the creation of the National Park several biomonitoring studies have been undertaken targeting different taxa, such as mammals (Ziegler 2002), fish (Crespi 1998), ungulates (Brugière 2005), birds (Nikolaus 1999), and amphibians and reptiles (Greenbaum & Carr 2005).  As detailed above several surveys on chimpanzees have been implemented (Ham 1998, Humle et al. 2009a, Fleury-Brugière & Brugière 2010, WCF 2012).
+
===Documented behaviours===  <!-- List of any behaviours observed at the site, including citations -->
In addition, studies were conducted that analysed the bushmeat trade in Haut Niger National Park (Brugière & Magassouba 2009, Humle & Alexandre 2015). Following the release of a group of rehabilitated chimpanzees their ranging patterns and behavior has been studied (Humle et al. 2009b, Humle et al. 2011).
        −
===Documented behaviours===  <!-- Paragraph describing any behaviours observed at the site, including citations -->
+
'''Table 6: Great ape behaviors reported for Haut Niger National Park'''
 +
{| border="1" cellpadding="5" cellspacing="0" class="behaviors-table"
 +
!align="left"|Behavior  <!-- Do not change categories -->
 +
!Source   <!-- source for behavior -->
 +
|-
 +
|Hand-clasp grooming
 +
|Humle et al. 2009b
 +
|-
 +
|}
   −
* hand-clasp grooming in chimpanzee group released in 2008 (Humle et al. 2009b)
      
===Relevant datasets===
 
===Relevant datasets===
 
[http://apesportal.eva.mpg.de/database/archiveMap A.P.E.S Portal]
 
[http://apesportal.eva.mpg.de/database/archiveMap A.P.E.S Portal]
      
<br>
 
<br>
    
= References =
 
= References =
[1] Bauer, H., De Iongh, H.H., Princée, F.P.G., & Ngantou, D. (2003) Research needs for lion conservation in West and Central Africa, C. R. Biologies, 326: S112–S118<br>
+
Bauer H et al. 2003. Research needs for lion conservation in West and Central Africa, C. R. Biologies, 326: S112–S118<br>
[2] BirdLife International (2018) Important Bird Areas factsheet: Mafou. Online: [http://datazone.birdlife.org/site/factsheet/mafou-iba-guinea/text  www.birdlife.org]<br>
+
BirdLife International. 2018. Important Bird Areas factsheet: Mafou. Online: [http://datazone.birdlife.org/site/factsheet/mafou-iba-guinea/text  www.birdlife.org]<br>
[3] Brugière, D. et al. (2005) Large- and medium-sized ungulates in the Haut Niger National Park, Republic of Guinea: population changes 1997–2002, Oryx, 39:50-55<br>
+
Brugière D et al. 2005. Large- and medium-sized ungulates in the Haut Niger National Park, Republic of Guinea: population changes 1997–2002, Oryx, 39:50-55<br>
[4] Brugière, D. et al. (2006) Population abundance of the common hippopotamus Hippopotamus amphibius in the Haut Niger National Park, Republic of Guinea, Mammalia:14-16<br>
+
Brugière D et al. 2006. Population abundance of the common hippopotamus Hippopotamus amphibius in the Haut Niger National Park, Republic of Guinea, Mammalia:14-16<br>
[5] Brugière, D. & Kormos, R. (2009) Review of the protected area network in Guinea, West Africa, and recommendations for new sites for biodiversity conservation, Biodiversity and Conservation, 18:847<br>
+
Brugière D, Kormos R. 2009. Review of the protected area network in Guinea, West Africa, and recommendations for new sites for biodiversity conservation, Biodiversity and Conservation, 18:847<br>
[6] Brugière, D. & Magassouba, B. (2009) Pattern and sustainability of the bushmeat trade in the Haut Niger National Park, Republic of Guinea, African Journal of Ecology, 44:630-639<br>
+
Brugière D, Magassouba B. 2009. Pattern and sustainability of the bushmeat trade in the Haut Niger National Park, Republic of Guinea, African Journal of Ecology, 44:630-639<br>
[7] Crespi, V. (1998) Preliminary study on the fishery resources of the River Niger in the Upper Niger National Park, Guinea, Fisheries Management and Ecology, 5:201–208<br>
+
Crespi V. 1998. Preliminary study on the fishery resources of the River Niger in the Upper Niger National Park, Guinea, Fisheries Management and Ecology, 5:201–208<br>
[8] Fleury-Brugière, M.-C. & Brugière, D. (2010) High Population Density of Pan troglodytes verus in the Haut Niger National Park, Republic of Guinea: Implications for Local and Regional Conservation, International Journal of Primatology, 31: 383-392<br>
+
Fleury-Brugière MC, Brugière D. 2010. High Population Density of Pan troglodytes verus in the Haut Niger National Park, Republic of Guinea: Implications for Local and Regional Conservation, International Journal of Primatology, 31: 383-392<br>
[9] Greenbaum, E. & Carr, J.L. (2005) The Herpetofauna of Upper Niger National Park, Guinea, West Africa, Natural History Museum The University of Kansas, 37:1-21 <br>
+
Greenbaum E, Carr JL. 2005. The Herpetofauna of Upper Niger National Park, Guinea, West Africa, Natural History Museum The University of Kansas, 37:1-21 <br>
[10] Ham (1998) Nationwide chimpanzee census and large mammal survey Republic of Guinea. <br>
+
Ham R. 1998. Nationwide chimpanzee census and large mammal survey Republic of Guinea.<br>
[11] Humle, T. et al (2009a) Report on chimpanzee surveys conducted in Parc National du Haut Niger, Republic of Guinea 2008-09<br>
+
Humle T et al. 2009a. Report on chimpanzee surveys conducted in Parc National du Haut Niger, Republic of Guinea 2008-09<br>
[12] Humle T, Colin, C., & Raballand, E. (2009b) Preliminary Report on Hand-Clasp Grooming in Sanctuary-Released Chimpanzees, Haut Niger National Park, Guinea, Pan Africa News, 16(1): 7-10<br>
+
Humle T et al. 2009b.  Preliminary Report on Hand-Clasp Grooming in Sanctuary-Released Chimpanzees, Haut Niger National Park, Guinea, Pan Africa News, 16(1): 7-10<br>
[13] Humle T, Colin, C., Laurans, M. & Raballand, E. (2011) Group Release of Sanctuary Chimpanzees (Pan troglodytes) in the Haut Niger National Park, Guinea, West Africa: Ranging Patterns and Lessons So Far, International Journal of Primatology, 32:456-473<br>
+
Humle T et al. 2011. Group Release of Sanctuary Chimpanzees (Pan troglodytes) in the Haut Niger National Park, Guinea, West Africa: Ranging Patterns and Lessons So Far, International Journal of Primatology, 32:456-473<br>
[14] Humle, T. & Konate, A. (2015) Primates and Bushmeat Hunting Around the High Niger National Park, Guinea, West Africa: Drivers and Patterns of Change, Folia Primatol, 86:235–386<br>
+
Humle T, Konate A. 2015. Primates and Bushmeat Hunting Around the High Niger National Park, Guinea, West Africa: Drivers and Patterns of Change, Folia Primatol, 86:235–386<br>
[15] Kormos, R. et al. (2003) The Republic of Guinea, in Kormos, R. Boesch, C., Bakarr, M.I. & Butynski, T.M. [eds.] Status Survey and Conservation Action Plan, IUCN/SSC Primate Specialist Group. IUCN, Gland, Switzerland and Cambridge, UK<br>
+
Kormos R et al. 2003. The Republic of Guinea, in Kormos, R. Boesch, C., Bakarr, M.I. & Butynski, T.M. [eds.] Status Survey and Conservation Action Plan, IUCN/SSC Primate Specialist Group. IUCN, Gland, Switzerland and Cambridge, UK<br>
[16] Kormos, R. & Boesch, C. (2003) Regional Action Plan for the Conservation of Chimpanzees in West Africa. IUCN/SSC Primate Specialist Group and Conservation International, Washington DC.<br>
+
Kormos R, Boesch C. 2003. Regional Action Plan for the Conservation of Chimpanzees in West Africa. IUCN/SSC Primate Specialist Group and Conservation International, Washington DC.<br>
[17] Lion Alert (2012) Online: [http://lionalert.org/alert/lions_in/Guinea  Lion Alert]<br>
+
Lion Alert. 2012. Online: [http://lionalert.org/alert/lions_in/Guinea  Lion Alert]<br>
[18] Nikolaus, G. (1999) The birds of the Parc National du Haut Niger, Guinea, Malimbus, 22, 1–22<br>
+
METT. 2009. Management Effectiveness Evaluations – Parc National du Haut Niger. Online: [https://papaco.org/guinea/  papaco.org]<br>
[19] Projet Primates (2016) Online: [https://www.projetprimates.com/en/  Chimpanzee Conservation Center (CCC)] <br>
+
Nikolaus G. 1999. The birds of the Parc National du Haut Niger, Guinea, Malimbus, 22, 1–22<br>
[20] Ramsar (2002Online: [https://rsis.ramsar.org/ris/1163  Ramsar 2002]<br>
+
Projet Primates. 2016. Online: [https://www.projetprimates.com/en/  Chimpanzee Conservation Center (CCC)] <br>
[21] Ramsar (2017) Online: [https://rsis.ramsar.org/ris/1164  Ramsar 2017]<br>
+
Ramsar. 2002. Online: [https://rsis.ramsar.org/ris/1163  Ramsar 2002]<br>
[22] UNESCO MAB (2011) UNESCO MAB Biosphere Reserve Directory. Haut Niger. Online: [http://www.unesco.org/new/en/natural-sciences/environment/ecological-sciences/biosphere-reserves/africa/guinea/haut-niger/  www.unesco.org]<br>
+
Ramsar. 2017. Online: [https://rsis.ramsar.org/ris/1164  Ramsar 2017]<br>
[23] WCF (2012) Etat de la faune et des menaces dans les aires protégées terrestres et principales zones de forte biodiversité de Rep. de Guinée. Report. Wild Chimpanzee Foundation, Sangaredi, Republic of Guinea. Online: [http://www.wildchimps.org Wild Chimpanzee Foundation]<br>
+
UNESCO MAB. 2011. UNESCO MAB Biosphere Reserve Directory. Haut Niger. Online: [http://www.unesco.org/new/en/natural-sciences/environment/ecological-sciences/biosphere-reserves/africa/guinea/haut-niger/  www.unesco.org]<br>
[24] Ziegler, S., Nikolaus, G. & Hutterer, R. (2002) High mammalian diversity in the newly established National Park of Upper Niger, Republic of Guinea, Oryx, 36:73-80<br>
+
WCF. 2012. Etat de la faune et des menaces dans les aires protégées terrestres et principales zones de forte biodiversité de Rep. de Guinée. Report. Wild Chimpanzee Foundation, Sangaredi, Republic of Guinea. Online: [http://www.wildchimps.org Wild Chimpanzee Foundation]<br>
 
+
Ziegler S et al. 2002. High mammalian diversity in the newly established National Park of Upper Niger, Republic of Guinea, Oryx, 36:73-80<br>
 
      
<br>
 
<br>
'''Page completed by:'''A.P.E.S. Wiki Team   '''Date:''' 06/12/18 <!-- If you don't want to add your name, you can add "A.P.E.S. Wiki team" -->
+
'''Page completed by:''' A.P.E.S. Wiki Team '''Date:''' 06/12/2018 <!-- If you don't want to add your name, you can add "A.P.E.S. Wiki team" -->
 
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