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   3) Keywords must be taken from the list of permitted words, or added to the list of permitted keywords. A link to the keyword list page is found adjacent  
 
   3) Keywords must be taken from the list of permitted words, or added to the list of permitted keywords. A link to the keyword list page is found adjacent  
 
     to the relevant table or keyword list. This ensures that the same keyword always refers to the same concept, e.g. Everybody uses "Industrial mining",  
 
     to the relevant table or keyword list. This ensures that the same keyword always refers to the same concept, e.g. Everybody uses "Industrial mining",  
     rather than several keywords such as "Industrial mining", "Large-scale mining", and "Mining".  
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     rather than several keywords such as "Industrial mining", "Large-scale mining", and "Mining". [[File: Map_CIV_MarahouéNP.png | 400px | thumb| right | (Côte d’Ivoire) Marahoué National Park © A.P.E.S. Wiki Team]]
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<!-- Please add information for this site -->
 
<!-- Please add information for this site -->
 
[[West Africa]] > [[Côte d’Ivoire]] > [[Marahoué National Park]]
 
[[West Africa]] > [[Côte d’Ivoire]] > [[Marahoué National Park]]
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[[File: Map_CIV_MarahouéNP.png | 400px | thumb| right | (Côte d’Ivoire) Marahoué National Park © A.P.E.S. Wiki Team]]
      
= Summary =  <!-- An overview of the site, with a one sentence overview of each of the following sections. can include a site map -->
 
= Summary =  <!-- An overview of the site, with a one sentence overview of each of the following sections. can include a site map -->
 
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<div style="float: right">
* Western chimpanzees ([https://www.iucnredlist.org/species/15935/17989872 ''Pan troglodytes verus'']) are now extirpated in Marahoué National Park.  
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{{#display_map: height=200px | width=300px | scrollzoom=off | zoom=5 | layers= OpenStreetMap, OpenTopoMap
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|7.11, -6.02~[[Marahoué National Park]]~Western Chimpanzee
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}}
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</div>
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* Western chimpanzees (''Pan troglodytes verus'') are now extirpated in Marahoué National Park.  
 
* This site has a total size of 1,010 km².  
 
* This site has a total size of 1,010 km².  
 
* Chimpanzees are extirpated due to hunting and large-scale deforestation driven by the expansion of agricultural areas.
 
* Chimpanzees are extirpated due to hunting and large-scale deforestation driven by the expansion of agricultural areas.
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Marahoué National Park (IUCN category: II) was established in 1968 (UNEP-WCMC and IUCN 2019). It covers an area of 1010 km2 and is located in the central Côte d´Ivoire in the Bouafle region near Lake Kossou.  Marahoué NP is 275 km (170 miles) northwest of Abidjan. Marahoué NP is managed by the Office Ivoirien des Parcs et Réserves (OIPR). The park consists of two ecological zones: close forest in the south-west of the park and savanna in the north-eastern part. In a survey in 2007 signs of different mammal and bird species were found, including forest elephant (''Loxodonta africana''), pygmy hippopotamus (''Choeropsis liberiensis''), African buffalo (''Syncerus caffer''), and bushbuck (''Tragelaphus scriptus''). However, the park is strongly affected by illegal activities, especially forest clearing for cocoa plantations and poaching, and the current status of the biodiversity is unknown (Campbell et al. 2008, N’Goran et al. 2006, N’Goran et al. 2007, WCF 2012).
 
Marahoué National Park (IUCN category: II) was established in 1968 (UNEP-WCMC and IUCN 2019). It covers an area of 1010 km2 and is located in the central Côte d´Ivoire in the Bouafle region near Lake Kossou.  Marahoué NP is 275 km (170 miles) northwest of Abidjan. Marahoué NP is managed by the Office Ivoirien des Parcs et Réserves (OIPR). The park consists of two ecological zones: close forest in the south-west of the park and savanna in the north-eastern part. In a survey in 2007 signs of different mammal and bird species were found, including forest elephant (''Loxodonta africana''), pygmy hippopotamus (''Choeropsis liberiensis''), African buffalo (''Syncerus caffer''), and bushbuck (''Tragelaphus scriptus''). However, the park is strongly affected by illegal activities, especially forest clearing for cocoa plantations and poaching, and the current status of the biodiversity is unknown (Campbell et al. 2008, N’Goran et al. 2006, N’Goran et al. 2007, WCF 2012).
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'''Table 1. Basic site information for Marahoué National Park'''
'''Table 1: Basic site information for Marahoué National Park'''
   
{| border="1" cellpadding="5" cellspacing="0" class="basic-information"
 
{| border="1" cellpadding="5" cellspacing="0" class="basic-information"
 
| Area            <!-- Please include units: km2/ha e.g 200ha    -->
 
| Area            <!-- Please include units: km2/ha e.g 200ha    -->
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|-
 
|-
 
| Coordinates
 
| Coordinates
|7.11 N, 6.02 W
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|7.11, -6.02
 
|-
 
|-
 
| Designation      <!-- National Park, Nature Reserve, etc.  -->
 
| Designation      <!-- National Park, Nature Reserve, etc.  -->
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|Subtropical/ tropical moist lowland forest, moist savanna, plantations
 
|Subtropical/ tropical moist lowland forest, moist savanna, plantations
 
|}
 
|}
[http://www.unitar.org/hiroshima/sites/unitar.org.hiroshima/files/Annex%201%20-%20IUCN%20Classification%20Schemes.pdf IUCN habitat categories]  [[Site designations]]
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[https://www.iucnredlist.org/resources/habitat-classification-scheme IUCN habitat categories]  [[Site designations]]
<br>
      
= Ape status =  <!-- a text overview of ape status (population sizes, trends etc), followed by a table of specific surveys and results -->
 
= Ape status =  <!-- a text overview of ape status (population sizes, trends etc), followed by a table of specific surveys and results -->
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The chimpanzee (''Pan troglodytes verus'') population in Marahoué National Park was first estimated at 1407 individuals by Marchesi et al. (1995). However, due to the relatively small number of transects surveyed, large uncertainties are associated with this estimate. In 2007, a survey that covered 167.5 km of systematically located transects across the entire park yielded a conservative population estimate of fewer than 50 individuals (N´Goran et al. 2007). In 2012, a rapid assessment was conducted by the Wild Chimpanzee Foundation [https://www.wildchimps.org  WCF] in collaboration with Afrique Nature, and no signs of chimpanzees were found.
 
The chimpanzee (''Pan troglodytes verus'') population in Marahoué National Park was first estimated at 1407 individuals by Marchesi et al. (1995). However, due to the relatively small number of transects surveyed, large uncertainties are associated with this estimate. In 2007, a survey that covered 167.5 km of systematically located transects across the entire park yielded a conservative population estimate of fewer than 50 individuals (N´Goran et al. 2007). In 2012, a rapid assessment was conducted by the Wild Chimpanzee Foundation [https://www.wildchimps.org  WCF] in collaboration with Afrique Nature, and no signs of chimpanzees were found.
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'''Table 2. Ape population estimates in Marahoué National Park'''
'''Table 2: Great ape population estimates in Marahoué National Park'''
   
{| border="1" cellpadding="5" cellspacing="0" class="population-estimate-table"
 
{| border="1" cellpadding="5" cellspacing="0" class="population-estimate-table"
 
! Species
 
! Species
 
! Year
 
! Year
! Abundance estimate (95% confidence interval)
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! Abundance estimate (95% CI)
! Density estimate (per km²)
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! Density estimate [ind./ km²] (95% CI)
 
! Encounter rate (nests/km)
 
! Encounter rate (nests/km)
 
! Area
 
! Area
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Marahoué National Park has lost 93% of its forest cover between 2002 and 2008, which may be directly linked to the civil unrest prevailing in the country since 2002 (Campbell et al. 2008). Deforestation was many driven clearing land for agriculture using fires, especially for cocoa plantations, and illegal logging (Campbell et al. 2008, N’Goran et al. 2006, N’Goran et al. 2007, WCF 2012). In addition, the presence of people in the park and illegal hunting of chimpanzees likely caused the extinction of chimpanzees in Marahoué NP (Struhsaker and Bakarr 1999, Denis 2015).
 
Marahoué National Park has lost 93% of its forest cover between 2002 and 2008, which may be directly linked to the civil unrest prevailing in the country since 2002 (Campbell et al. 2008). Deforestation was many driven clearing land for agriculture using fires, especially for cocoa plantations, and illegal logging (Campbell et al. 2008, N’Goran et al. 2006, N’Goran et al. 2007, WCF 2012). In addition, the presence of people in the park and illegal hunting of chimpanzees likely caused the extinction of chimpanzees in Marahoué NP (Struhsaker and Bakarr 1999, Denis 2015).
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'''Table 3. Threats to great apes in Marahoué National Park'''
'''Table 3: Threats to great apes in Marahoué National Park'''
   
{| border="1" cellpadding="5" cellspacing="0" class="threats-table"
 
{| border="1" cellpadding="5" cellspacing="0" class="threats-table"
 
!align="left"|Category  <!-- Do not change threat categories -->
 
!align="left"|Category  <!-- Do not change threat categories -->
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|-
 
|-
 
|1. Residential & commercial development
 
|1. Residential & commercial development
|1.1. Housing & urban areas
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|1.1 Housing & urban areas
 
|High
 
|High
 
|
 
|
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|-
 
|-
 
|2. Agriculture & aquaculture
 
|2. Agriculture & aquaculture
|2.1. Annual & Perennial Non-Timber Crops
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|2.1 Annual & Perennial Non-Timber Crops
 
|High
 
|High
 
|
 
|
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|-
 
|-
 
|
 
|
|2.3. Livestock Farming & Ranching
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|2.3 Livestock Farming & Ranching
 
|Low
 
|Low
 
|
 
|
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|-
 
|-
 
|4. Transportation & service corridors
 
|4. Transportation & service corridors
|4.1. Roads & railroads
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|4.1 Roads & railroads
 
|Medium
 
|Medium
 
|
 
|
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|-
 
|-
 
|5. Biological resource use
 
|5. Biological resource use
|5.1. Hunting & collecting terrestrial animals
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|5.1 Hunting & collecting terrestrial animals
 
|High
 
|High
 
|
 
|
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As chimpanzees are extirpated from Marahoué National Park no conservation activities are implemented.
 
As chimpanzees are extirpated from Marahoué National Park no conservation activities are implemented.
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'''Table 4. Conservation activities in Marahoué National Park'''
'''Table 4: Conservation activities in Marahoué National Park'''
   
{| border="1" cellpadding="5" cellspacing="0" class="conservation-actions-table"
 
{| border="1" cellpadding="5" cellspacing="0" class="conservation-actions-table"
 
!align="left"|Category  <!-- Do not change threat categories -->
 
!align="left"|Category  <!-- Do not change threat categories -->
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[[Conservation activities list (Junker et al. 2017)]]
 
[[Conservation activities list (Junker et al. 2017)]]
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= Impediments = <!-- Overview of impediments to ape conservation -->
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= Challenges = <!-- Overview of impediments to ape conservation -->
    
Denis (2015) conducted a comprehensive study into why Marahoué National Park degraded.  The study was based on interviews with different stakeholders in Marahoué National Park and pointed out that already since its creation in 1968 farmers were present in the park and in 1990 13% of the park was occupied by people, and this percentage has grown steadily since (Denis 2015). Government support for maintaining the protective status of Marahoué was weak and inconsistent. There is also widespread corruption and a lack of technical means as well as human resources to ensure law enforcement (Denis 2015).
 
Denis (2015) conducted a comprehensive study into why Marahoué National Park degraded.  The study was based on interviews with different stakeholders in Marahoué National Park and pointed out that already since its creation in 1968 farmers were present in the park and in 1990 13% of the park was occupied by people, and this percentage has grown steadily since (Denis 2015). Government support for maintaining the protective status of Marahoué was weak and inconsistent. There is also widespread corruption and a lack of technical means as well as human resources to ensure law enforcement (Denis 2015).
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'''Table 5. Challenges reported for Marahoué National Park'''
'''Table 5: Impediments reported for Marahoué National Park'''
   
{| border="1" cellpadding="5" cellspacing="0" class="impediments-table"
 
{| border="1" cellpadding="5" cellspacing="0" class="impediments-table"
!align="left"|Impediment <!-- Do not change categories -->
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!align="left"|Challenge <!-- Do not change categories -->
 
!Source  <!-- source for impediment mentioned -->
 
!Source  <!-- source for impediment mentioned -->
 
|-
 
|-
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'''Table 6: Great ape behaviors reported for Marahoué National Park'''
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'''Table 6. Great ape behaviors reported for Marahoué National Park'''
 
{| border="1" cellpadding="5" cellspacing="0" class="behaviors-table"
 
{| border="1" cellpadding="5" cellspacing="0" class="behaviors-table"
 
!align="left"|Behavior  <!-- Do not change categories -->
 
!align="left"|Behavior  <!-- Do not change categories -->
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===Relevant datasets===
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[http://apesportal.eva.mpg.de/database/archiveMap A.P.E.S Portal]
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  −
<br>
      
= References =
 
= References =

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